Thirteen QTLs for grain protein content were identified in a RI population from the cross WL711 (low protein content)/PH132 (high grain content) {10055}. The QTLs that were identified using more than one method or in more than one environment are listed below. Also listed is a QTL that was identified in the mean over the four environments and was therefore deemed important {10055}. QTLs for grain protein content were detected on chromosome arms 6AS (associated AFLP marker, XE38M90v200v ) and 1BL (associated RFLP marker, Xcdo1188 ) in Courtot/Chinese Spring’ {0141}.

QGpc.ndsu-6Ba

Chromosome	6B
Type varieties	Langdon {623}.

QGpc.ndsu-6Bb {10071}, {623}. 6B.

GPC-B1

Gpc-B1a

Synonym

QGpc.ndsu-6Ba {623}

This allele, fixed in cultivated durum, is a non-functional frame-shift mutation {10438}. A similar nonfunctional allele, or a complete deletion of GPC-B1 , is fixed in hexaploid wheat {10438}.

Gpc-B1b

Synonym	QGpc.ndsu-6Bb {10071}, {623}, Gpc-6B1 {10299}, NAM-B1 {10995}
Chromosome	6BS
i	Yecora Rojo NIL PI 638740 {10138}.
Varieties	As II {10995}; Burnside {11044}; Diamant {10995}; Glencross {11044}; Glupro {10138}; Lilian {11044}; Prins {10995}; Somerset {11044}; Stanley
Type varieties	T. dicoccoides FA-15 {10138}.
Marker associations	Mapped to a 0.3 cM interval flanked by Xucw79-6B and Xucw71-6B {10229}; Xcdo365-6B – 1.5 cM – Gpc-B1 – 1.2 cM – Xucw67-6B {10296.

Gpc-B1b , the functional allele {10438} in T. dicoccoides , affects senescence and maturity in addition to grain protein content, accelerating senescence and maturity {10298}. Gpc-B1 is a NAC transcription factor designated Nam-B1 {10438}. A paralogous copy of this gene is present in homologous group 2 ( Nam2 ). This allele was relatively frequent in Scandinavian and Finnish common wheats, landraces and spelts {10995}.

PRO1

Chromosome	5DL
Sources / synonyms	CS[] 6/Hope 5D {777}. May be identical to VRN-D1* .

PRO2

Chromosome	5DS
Sources / synonyms	CS[*] 6/Hope 5D {777}.

QGpc.ccsu-2B.1

Chromosome	2BL
Varieties	WL711/PH132 RI mapping population {10055}; higher protein content was contributed by PH132 and the QTL explained 13.4% of the phenotypic variation {10055}.
Marker associations	Associated with Xgwm1249-2B {10055}.

QGpc.ccsu-2D.1

Chromosome	2DL
Varieties	WL711/PH132 RI mapping population {10055, 0015}; higher protein was contributed by PH132 and the QTL explained 19% {0015} and 14% {10055} of the phenotypic variation.
Marker associations	Associated with Xgwm1264-2D {10055}.

QGpc.ccsu-3D.1

Chromosome	3DS
Varieties	WL711/PH132 RI mapping population {10055}; higher protein content was contributed by PH132 and the QTL explained 16.3% of the phenotypic variation {10055}.
Marker associations	Associated with Xgwm456-3D {10055}.

QGpc.ccsu-3D.2

Chromosome	3DS
Varieties	WL711/PH132 RI mapping population {10055}; higher protein content was contributed by PH132 and the QTL explained 14% of the phenotypic variation {10055}.
Marker associations	Associated with Xgwm892-3D {10055}.

QGpc.ccsu-7A.1

Chromosome	7AS
Varieties	WL711/PH132 RI mapping population {10055}; higher protein content was contributed by PH132 and the QTL explained 32.4% of the phenotypic variation {10055}.
Marker associations	Associated with Xgwm1171-7A {10055}.

QGpc.ipk.7B

Varieties	F26-70 {10628}; Closely associated with Ppd-B2 {10628}.
su	Favorit (F26-70 7B) {10628}. See Response to Photoperiod.

QGpc.ndsu-5B.1

Chromosome	5B
Varieties	LDN (DIC5B)/LDN, contributed by DIC5B {10161}.
Marker associations	Nearest marker, Xgwm604-5B {10161}.

QGpc.ndsu-5B.2

Chromosome	5B
Varieties	LDN (DIC5B)/LDN, contributed by DIC5B {10161}.
Marker associations	Nearest marker, Xabc310-5B {10161}.

QGpc.ndsu-5B.3

Chromosome	5B
Varieties	LDN (DIC5B)/LDN, contributed by DIC5B {10161}.
Marker associations	Nearest marker, Xwg909-5B {10161}.

QGpc.ndsu-6B

Chromosome	6BS
Type varieties	Langdon {623}.

QPro.inra-2A

Chromosome	2A
Varieties	Renan/Recital {10071}.
Marker associations	XksuD18-2A – Xgwm614-2A (R[2 ] = 4.4-8.9%) {10071}.

QPro.inra-3A

Chromosome	3A
Varieties	Renan/Recital {10071}.
Marker associations	Xcfd79-3A – Xfbb250-3A (R[2 ] = 4.1-8.3%) {10071}.

QPro.inra-4D

Chromosome	4D
Varieties	Renan/Recital {10071}.
Marker associations	Linked to Xcfd71-4D (R[2 ] = 4.610.3%) {10071}.

QPro.inra-7D

Chromosome	7D
Varieties	Renan/Recital {10071}.
Marker associations	Xcfd69-7D – Pch1 (R[2] =6.410.4%) {10071}.

QPro.mgb-4B

Associated with Gai1 and Xpsr622-4B {110}[2] .

QPro.mgb-5A

Associated with Xpsr911-5A {110}[2] and Xcdo412-5A {0343}[*]

QPro.mgb-6A.1

Associated with Xpsr167-6A and XksuG8-6A {110}[2] .

QPro.mgb-6A.2

Associated with Xmgb56-6A {110}[2] and Xpsr627-6A {0343}[*] .

QPro.mgb-6B

Note	Associated with Gli-B2-6B {110}[2] and Nor-2 {0343}[*] .
Marker associations	QGpc.ndsu-6B was associated (LOD score =18.9) with the interval Xmwg79-6B – Xabg387-6B . These loci were mapped in 6BS: Xmwg79-6B – 5.9 cM – Xabg387-6B

Qpro.mgb-7A

Associated at P<=0.01 with Pan2 {0343}[*] .

QPro.mgb-7B

Associated with Xpsr490(Ss1)-7B , Pc {110}[2] and Xutv913-7B {0343}[*] . QTLs for grain protein content were detected on chromosome arms 6AS (associated AFLP marker, XE38M60 200) and 1BL (associated RFLP marker, Xcdo1188-1B ) in Courtot/Chinese Spring {0141}. Forno / Oberkulmer spelt: Nine QTLs (51% of the variation) were mapped in cross {0280}. Cheyenne (high quality wheat) / CS (low quality wheat): RSL population: A QTL for grain and flour protein content, contributed by CS, was associated with XTri-1D /Centromere {0251}. Renan / Recital: Four QTL conferring grain protein content {10071}; only QTL stable over at least 4 of 6 locations were presented. Renan contributed the four alleles for high grain protein content. Ning 7840 / Clark: RIL population: QTL from Ning 7840 were detected on chromosomes 3AS ( Xwmc749-3AS – Xgwm369-3AS ; R[2 ] = 0.09-0.11) and 4B ( Xgwm368-4B – Xwmc617-4B , R[2 ] = 0.08-0.11) {10702}. ## Tetraploid wheat T. dicoccoides / Latino: In line 3BIL-85 high grain protein was detected in chromosomes 2AS (associated with Xcfa2164-2A , R[2 ] = 17%), 6AS ( Xp39M37 250-6A, R[2 ] = 17%) and 7BL ( Xgwm577-7B , R[2 ] = 9%) {10338}.

2.2. Enzymes⌂ Home

ACPH-1

Acph-A1

Synonym	Acph-B1 {936}, Acph3 {516}, Acph2 {516}
Chromosome	4AS
Varieties	CS.

Acph-B1

Synonym

Acph8 {516}, Acph4 {516}, Acph-A1 {936}

4BL {504}, {516}.

Acph-D1

Synonym

Acph5 {516}, Acph6 {516}

4DL {504}, {516}.

Acph-H1

Chromosome	4H
ad	CS/Betzes. Acph-M[v] 1 {237}. [ Acph-M[v] 1 {985}, Aph-v {237}]. 4M[v ] {237}.
tr	H-93-33 {984}.

Acph-R1

Chromosome	7R
Chromosome	7RS
ad	CS/Imperial. Acph-S[s] 1 {1140}. 4S[s ] {1140}.
ad	CS/ T. searsii .

ACPH-2

Acph-D2

Synonym	Acph1 {10309}
Chromosome	2DL
dv	Acph-D2 100 and Acph-D2 95 alleles distinguished accessions of Ae. tauschii ssp. tauschii and strangulata , respectively {1030}.
Type varieties	Ae. tauschii {10407}.
Marker associations	Cent ... Acph-D2 – 4 cM – Xgwm157- 2D {10309}.

Acid phosphatase gene loci were reported for 7RL in S. cereale {1251}, chromosomes L1 (= 7Ag[i] ) and L4 (= 4Ag[i] ) of Thin. intermedium {361}, and chromosome E of Ae. umbellulata {0069}. Two loci on 7R were separated by 25 +or- 5.2 cM {1534}. Wehling {1559} identified four acid phosphatase loci in S. cereale , three of which were in 7R.

ADH-A1

Synonym	Adh-B1 {504}, AdhB {502}
Chromosome	4A

4AL {504}, {516}.

Adh-B1a

Synonym	Adh11 {501}, Adh-A1a {1442}
Varieties	CS.
Type varieties	PI 226951 {501}; Malavika {1442}.

Adh-B1b

Synonym	Adh12 {501}, Adh-A1b {1442}
Varieties	Rageni derivative {1443}.
Type varieties	CI 4013 {501}; Bijaga Yellow {1442}. Adh-B1b was the only variant ADH-1 allele detected in study of a large number of T. aestivum and _T.

ADH-D1

Synonym	AdhD {502}
Chromosome	4D
Chromosome	4DS
Varieties	CS.
Marker associations	Adh-D1 [ Adh1, Adh2 ] was mapped 4 cM distal to Xpsr163-4D and closely proximal to Xcsiha114-4D.1 . [ XcsIHA114-1a '] {757}.

ADH-C1

Synonym	G {1278}
ad	T. aestivum cv. Alcedo/ Ae. caudata line G. ADH-Ag[i] 1 {560}, {374}. [ Adh-X1 {361}]. 4Ag[i] {560}.
ad	Vilmorin 27/ Th. intermedium ; Caribo/ Th. intermedium .

ADH-E1

Chromosome	4ES
ad	CS/ E. elongata .

ADHG-H1

Chromosome	4H
ad	CS/Betzes. ADH-M[v] 1 {984}. [ ADHmu {984}, Adh-M[v] 1 {985}]. 4M[v ] {984}.
Varieties	H-93-33.

ADH-R1

Synonym	AdhR2 {582}
Chromosome	4R
Chromosome	4RS
ad	CS/Imperial {1457, 506}; FEC28/Petkus {43}; Holdfast/King II {582}. ADH-V1 {1026}, {242}. 4V {1026}.
ad	CS/ D. villosum .

Three Adh genes were identified in Hordeum vulgare and H. spontaneum {144}, {490}, {493}, {520}. Two of these were tightly linked at the Adh-H1 locus {144}. The third gene was tentatively located in 5H {490}. A low-level of aliphatic alcohol dehydrogenase activity is commonly observed on zymograms in the absence of added substrate {513}; this may account for the observation of wheat lactate dehydrogenase that was reported in {1465}. The gene series formerly designated Adh-2 and Adh-3 appear under Aromatic Alcohol Dehydrogenase

AMP-A1

Note	6AS {504}, {516}.
Varieties	CS.

Amp-A1a

Varieties

CS {1533}.

Amp-A1b

Varieties

Vitka {1533}.

AMP-B1

Note	6BS {504}, {516}.
Varieties	CS.

Amp-B1a

Varieties

CS {1533}.

Amp-B1b

Varieties	Iskra {1533}. Amp-B1c {703}, {1244}. Null allele.
Varieties	T. spelta IPSR 1220017 {703}; Sinvalocho M.A {1244}.

AMP-D1

Note	6DS {504}, {516}.
Varieties	CS.

Amp-D1a

Varieties

CS.

Amp-D1b

Varieties

Sears' Synthetic IPSR1190903.

AMP-Ag[e] 1 {1575}. 6Ag[e] {1575}. ad,su: Rescue/ Th. elongatum . AMP-Ag[i] 1 {703}. 6Ag[i] {703}.

AMP-C1

Chromosome	6D
ad	Alcedo/ Ae. caudata line D.

AMP-E1

Chromosome	6E
ad	CS/ E. elongata .

AMP-H1

Chromosome	6H
ad	CS/Betzes.

AMPp-R1

Chromosome	6R
ad	CS/Imperial {1457}; Holdfast/King II {1280}.

a-Amy1c

Synonym	a-Amy-B4
Type varieties	T. durum ssp. georgicum . The presence of a-Amy1 reported in {1084} was confirmed by tests of segregation in a CS/Jones Fife population and in

A further set of a-amylase genes, Xa-Amy-5 [ a-Amy3 ], was identified in 5A, 5B and 5D by crosshybridization with a-AMY-1 and a-AMY-2 probes {80}. Only one gene copy appears to be present at each locus. In rye, evidence was obtained for three a-Amy-1 genes, two or three a-Amy-2 genes and three aAmy-3 genes {907}. Synthesis of a-amylase isozymes controlled by a-Amy-1 genes on chromosomes 6A and 6D is reduced in DT6BS compared to euploid CS. This result suggests the presence of a gene(s) on the long arm of chromosome 6B, which is (are) required for GA-induced alpha-amylase synthesis in the aleurone {0072}.

EP-D1

Chromosome	7DL
Varieties	CS.

Ep-D1a

Varieties

CS.

Ep-D1b

Synonym	EP-V1 {973}
Varieties	5L 219 {1521}; H-93-70 {1521}; Hyak {21}; Madsen {20}; Rendezvous {708}; VPM1 {973}. Assuming that Ep-D1 encoded an oligopeptidase G,

Ep-D1c

Varieties

Sears' Synthetic.

Ep-D1d

Note	Null allele.
Varieties	Wheats with Lr19 {1587}.

Ep-D1e

Note	Isozyme 5.
Varieties	PI 294994 {894}.

EP-E1

Chromosome	7EL
al	CS/ E. elongata .

EP-H1

Chromosome	7HL
al	CS/Betzes.

EP-H[ch] 1

Chromosome	7H
su	CS/ H. chilense .

EP-H[t] 1

Note	7H[t] S {1037}.
ad	CS/ E. trachycaulus .

EP-M[v] 1

Synonym	Ep-M[v] 1 {985}
Chromosome	7M

EP-R1 {92}, {708}, {266}. 6RL {92}.

EP-S[b] 1

Chromosome	7S
su	Holdfast/ Ae. bicornis .

EP-S[l] 1

Chromosome	4S
ad	CS/ Ae. longissima .

EP-S[s] 1

Chromosome	7S
ad	CS/ T. searsii .

EP-U1

Chromosome	7U
su	CS/ Ae. umbellulata .

EP-V1

Chromosome	7V
ad	CS/ D. villosum .

GPI-Ag[i] 1

Synonym

Gpi-X1 {361}

1Ag[i ] {361}.

GPI-E1

Chromosome	1ES
ad	CS/ E. elongata .

GPI-H1

Chromosome	1HS
ad	CS/Betzes.

GPI-H[ch] 1

Chromosome	1H
ad	CS/ H. chilense .

GPI-R1

Chromosome	1R
Chromosome	1RS
ad	CS/King II {195}.
al	2a, 2b, and R14 {779}.

GPI-R[m] 1

Chromosome	1R
ad	CS/ S. montanum .

GPI-S[l] 1

Note	1S[1] S {1228}.1S[1] {517}.
Marker associations	In Ae. longissima 2 x Ae. longissima 10, GPI-S[l] 1 , two glutenin loci, and three gliadin loci were mapped relative to one another as

GPI-S[s] 1

Chromosome	1S
ad	CS/ Ae. searsii .

GPI-U1

Chromosome	1U
ad	CS/ Ae. umbellulata .

GPI-V1

Chromosome	1V
ad	CS/ D. villosum .

GOT-A2

Chromosome	6AL
Varieties	CS.

GOT-B2

Chromosome	6BL
Varieties	CS.

GOT-D2

Chromosome	6DL
Varieties	CS.
Marker associations	Cent – Got-D2 – 2 cM – Xpsr154-6D {757}.

GOT-Ag[e] 2

6Ag[e ] {1575}. ad,su: Rescue/ Th. elongatum .

GOT-E2

Note	6EBeta {518}.
ad	CS/ E. elongata .

GOT-H2

Chromosome	6H
ad	CS/Betzes.

GOT-H[t] 2

Chromosome	6H
ad	CS/ E. trachycaulus .

GOT-R2

Synonym

Got3 {1559}

6R {1457}; 6RL {1280}.

GOT-3 GOT-A3

Chromosome	3AL
Varieties	CS.

GOT-B3

Chromosome	3BL
Varieties	CS.

GOT-D3

Chromosome	3DL
Varieties	CS.

GOT-Ag[e] 3

Note	3Ag[e] L {521}.
ad	CS/TAP 67.
su	CS/TAP 67.
tr	Certain CS 3D/Ag lines.

GOT-C3

Note	F {1278}.
ad	T. aestivum cv. Alcedo / Ae. caudata line C.

Got-E3

Chromosome	3EL
ad	CS/ E. elongata .

GOT-H3

Synonym	Got-b3 {90}
Chromosome	3H
ad	CS/Betzes.

GOT-H[ch] 3

Chromosome	3H
ad	MA/ H. chilense .

GOT-R3

Synonym	Got3 {1559}
Chromosome	3R
ad	CS/Imperial {1457}; Holdfast/ King II {1253}; Kharkov/Dakold {1253}.

GOT-S[s] 3

Chromosome	3S
ad	CS/ Ae. searsii . GOT-V3 {1518}, {242}. 3VL {1518}.
ad	Creso/ D. villosum . GOT-4

GOT-R4

Synonym	Got1/7R {1203}, Got2 {1559}
Chromosome	7RL
al	S. cereale .

Mdh-A3b

Varieties

Bersee.

MDH-B1

Synonym	Mdh2B {87}
Chromosome	1BL

Mdh-B3a

Varieties

CS.

Mdh-B3b

Varieties

Hope.

MDH-D1

Synonym	Mdh2D {87}
Chromosome	1DL
Varieties	CS.

MDH-H1

Chromosome	1HL
ad	CS/Betzes.

MDH-H[ch] 1

Chromosome	1H
ad	MA/ H. chilense .

MDH-R1

Synonym	Mdh2-1 {1252}
Chromosome	1RL
ad	CS/Imperial 1R; Kharkov/Dakold 1R; Holdfast/King II 1RL.

MDH-S[s] 1

Chromosome	1S
ad	CS/ T. searsii .

MDH-2

MDH-H2

Synonym	Mdh2-b2 {90}
Chromosome	3H

MDH-R2

Synonym	Mdh2-2 {1252}
Chromosome	3R
ad	CS/Imperial.

A third set of dimeric MDH isozymes identified in mature grain was separable from MDH-1 and MDH-2 by their higher pI's in IEF {811}.

MDH-3 MDH-A3

Chromosome	5AS
Varieties	CS.

Mdh-D3a

Varieties

CS.

Mdh-D3b

Varieties

Sears' Synthetic.

MDH-B3

Chromosome	5BS
Varieties	CS.

MDH-D3

Chromosome	5DS
Varieties	CS.

MDH-E3

Chromosome	5ES
ad	CS/ E. elongata .

MDH-H3

Chromosome	5H
ad	CS/Betzes.

MDH-U3

Chromosome	5U
ad	CS/ Ae. umbellulata .

MDH-R4

Chromosome	1RL
Varieties	Various crosses.

PER-B1

Chromosome	1BS
Varieties	CS.

PER-D1

Chromosome	1DS
Varieties	CS.

Per-D1a

Varieties

CS.

Per-D1b

Varieties

Sears' Synthetic.

PER-H[ch] 1

Chromosome	1H
ad	CS/ H. chilense .

PER-R1

Synonym	Prx {1561}
Chromosome	1RS
ad	CS/King II {12}; Holdfast/King II {1561}.
tr	Veery 'S' {12}.

PER-V1

Chromosome	1V
ad	Creso/ D. villosum .

PER-2 . PER-2 is expressed in young leaf {118}, coleoptile and root {816} tissues.

PER-A2

Chromosome	2AS
Varieties	CS.

Per-A2a

Varieties

CS.

Per-A2b

Varieties

Timstein.

PER-B2

Chromosome	2BS
Varieties	CS.

Per-B2a

Varieties

CS.

Per-B2b

Varieties

Sears' Synthetic IPSR1190903.

PER-D2

Chromosome	2DS
Varieties	CS.

PER-H2

Synonym	Per-5 {95}
Chromosome	2H
ad	CS/Betzes.

PER-R2

Chromosome	2RS
ad	CS/Imperial; Kharkov/Dakold. PER-3. PER-3 is expressed in embryo {119, 816} and scuteller {119} tissues.

PER-A3

Chromosome	3AL
Varieties	CS.

Per-A3a

Varieties

CS.

Per-A3b

Varieties

Timstein.

Per-A3c

Varieties

Hobbit 'S'.

PER-B3

Synonym	Per4 {961}
Chromosome	3BL
Varieties	CS.

Per-B3a

Varieties

CS.

Per-B3b

Varieties

Hope.

Per-B3c

Varieties

T. macha IPSR1240005.

Per-B3d

Varieties

Timstein.

Per-B3e

Varieties

Sears' Synthetic IPSR1190903.

PER-D3

Synonym	Per5 {961}
Chromosome	3DL
Varieties	CS.

Per-D3a

Varieties

CS.

Per-D3b

Varieties

Hope.

Per-D3c

Varieties

Timstein.

Per-D3d

Varieties

T. macha IPSR 142005.

Per-D3e

Varieties

Sava.

Per-D3f

Varieties

Cheyenne.

Per-D3g

Varieties

Sears' Synthetic IPSR 1190903. Varietal variation for PER-3 was reported in {94}. PER-4 . PER-4 is expressed in endosperm tissue {86, 119}.

PER-A4

Synonym	Per3 {961}
Chromosome	7A
Chromosome	7AS
Varieties	CS.

Per-A4a

Varieties

CS.

Per-A4b

Varieties

Hope.

Per-A4c

Varieties

Sicco

PER-B4

Synonym	Per2 {961}
Chromosome	4A
Chromosome	4AL
Varieties	CS.

Per-B4a

Varieties

CS.

Per-B4b

Varieties

Hope.

Per-B4c

Varieties

Thatcher.

PER-D4

Synonym

Per1 {961}

7D {695},7DS {694, 86, 119}.

Per-D4a

Varieties

CS.

Per-D4b

Varieties

Thatcher.

PER-Ag[e] 4

Note	7Ag[e] S {694}.
tr	Certain CS 7D/ Ag[e] lines.

PER-Ag[i] 4

Synonym

Per-Ag[i] 3 {374}

7Ag[i] {168}.

PER-D5

Chromosome	2DS
Varieties	CS.

PER-S[l] 5

Chromosome	2S
ad	CS/ Ae. longissima

PDE-A1

Synonym	Pde-A3 {1590}
Chromosome	3AS
Chromosome	3A
Varieties	CS.

PDE-B1

Synonym	Pde-B3 {1590}
Chromosome	3BS
Chromosome	3B
Varieties	CS.

PDE-D1

Synonym	Pde-D3 {1590}
Chromosome	3DS
Varieties	CS.

PDE-S[l] 1

Note	3S[l] S {172}.
ad	CS/ Ae. longissima .

PDE-V1

Chromosome	3VS
ad	CS/ D. villosum .

PGD1

Synonym

Pgd-A3 {963}, Pgd3 {282}

7A[m] S {282}.

PGDR1

Chromosome	4RL
ad	CS/Imperial; Holdfast/King II.

PGDR2

Chromosome	6RL
ad	CS/Imperial; Holdfast/King II. Loci were also identified in 6B {1435}, 1EL {1435}, 1HL {147}, {1072}, 1H[ch] {352} and 1RL {779}.

PGM-A1

Synonym	Pgm-B1 {88}
Chromosome	4AL
Varieties	CS.

PGM-D1

Chromosome	4DS
Varieties	CS.

PGM-Ag[i] 1

Synonym

Pgm-X1 {361}

4Ag[i ] {361}.

PGM-H1

Synonym	Pgm-b1 {90}
Chromosome	4H
ad	CS/ Betzes.

PGM-H[ch] 1

Chromosome	4H
ad	MA/ H. chilense .

PGM-R1

Chromosome	4RS
ad	CS/Imperial 4RS {1561, 1253}; Kharkov/Dakold 4R {1253}; Holdfast/King II 4RS{1561, 1253}.

SKDH-H1

Chromosome	5H
ad	CS/Betzes.

SKDH-H[t] 1

Chromosome	5H
ad	CS/ E. trachycaulus .

SKDH-M[v] 1

Synonym

Skdh-M[v] 1 {985}

5M[v] .

SKDH-R1

Chromosome	5RS
Chromosome	5R
ad	CS/King II {85}; CS/Imperial {706}; Kharkov/Dakold {85}.
tr	CS 4AS-5RL; CS 5BL-5RL.

SKDH-S[l] 1

Note	5S[l] S {85}.
ad	CS/ Ae. longissima .

SKDH-S[s] 1

Chromosome	5S
ad	CS/ Ae. searsii .

SKDH-U1

Chromosome

ad,su: CS/ Ae. umbellulata.

SKDH-V1

Chromosome	5V
ad	CS/ D. villosum .

SOD-1 SOD-A1

Chromosome	2AL
Varieties	CS.

SOD-B1

Chromosome	2BL
Varieties	CS.

SOD-D1

Chromosome	2DL
Varieties	CS.

SOD-E1

Note	VI E {808}.
ad	CS/ E. elongata .

SOD-H1

Chromosome	2H
ad	CS/Betzes.

SOD-R1

Synonym	Sod-3 {586}
Chromosome	2R
ad	CS/Imperial.

SOD-S[s] 1

Chromosome	2S
ad	CS/ Ae. searsii .

SOD-VI

Chromosome	7V
ad	CS/ D. villosum .

TPI-1 TPI-A1

Chromosome	3AS
Varieties	CS.

TPI-B1

Chromosome	3BS
Varieties	CS.

TPI-D1

Chromosome	3DS
Varieties	CS.

TPI-E1

Chromosome	3E
ad	CS/ E. elongata .

TPI-H1

Chromosome	3H
ad	CS/Betzes.

TPI-R1

Chromosome	3R
ad	CS/Imperial; Kharkov/Dakold.

TPI-S[l] 1

Chromosome	3S
ad	CS/ Ae. longissima .

TPI-2 TPI-A2

Chromosome	5AL
Varieties	CS.

TPI-B2

Chromosome	5BL
Varieties	CS.

TPI-D2

Chromosome	5DL
Varieties	CS.

TPI-Ag[i] 2

Note	5Ag[i ] {374}.
ad	Vilmorin 27/ Th. intermedium .

TPI-H2

Chromosome	5H
ad	CS/Betzes.

TPI-R2

Chromosome	5R
ad	CS/Imperial; Kharkov/Dakold.

TPI-S[l] 2

Chromosome	5S
ad	CS/ Ae. longissima .

TPI-U2

Chromosome	5U
ad	CS/ Ae. umbellulata .

ACO-A1

Chromosome	6AL
Varieties	CS.

Aco-A1a

Varieties

CS {1533}.

Aco-A1b

Varieties

Dubravka {1533}.

ACO-B1

Chromosome	6BL
Varieties	CS.

Aco-B1a

Varieties

CS {1533}.

Aco-B1b

Varieties

Dubravka {1533}.

Aco-B1c

Varieties

Slavonka {1533}.

ACO-Ag[e] 1

6Ag[e ] {1575}. ad,su: Rescue/ Th. elongatum .

ACO-E1

Note	6Ebeta {189}.
ad	CS/ E. elongata .

ACO-H1

Synonym	Aco-1 {147}
Chromosome	6HL
Chromosome	6H
ad	CS/Betzes.

ACO-R1

Chromosome	6RL
ad	Sturdy/PI 252003.

ACO-S[l] 1

Chromosome	6S
ad	CS/ Ae. longissima .

ACO-S[s] 1

Chromosome	6S
ad	CS/ Ae. searsii .

ACO-U1

Note	CSU-31 {189}.
ad	CS/ Ae. umbellulata .

ACO-2 ACO-B2 Aco-B2a

Varieties

CS.

Aco-B2b

Varieties

PI 278437.

Aco-B2c

Varieties

PI 182575.

Aco-B2d

Varieties

PI 157589.

ACO-D1

Chromosome	6DL
Varieties	CS.

Further alleles at Aco-A1 and Aco-B1 are listed in {1127}; these have not been tested against those found in {1533}.

ACO-A2

Chromosome	5AL
Varieties	CS.

ACO-B2

Chromosome	4BL
Varieties	CS.

ACO-D2

Chromosome	4DL
Varieties	CS.

ACO-E2

Chromosome	4EL
ad	CS/ E. elongata .

ACO-M[v] 2

Synonym

Aco-M[v] 2 {985}

5M[v] .

ACO-R2

Chromosome	5RL
ad	CS/King II 5R; Holdfast/ King II 5RL.

ACO-S[s] 2

Chromosome	4S
ad	CS/ Ae. searsii .

NDH-1 NDH-A1

Synonym	Ndh-B1 {513}
Chromosome	4AL
Varieties	CS.

Ndh-A1a

Synonym	Ndh-B1a {936}
Varieties	CS.

Ndh-A1b

Synonym	Ndh-B1b {936}
Varieties	Sutjeska.

Ndh-A1c

Synonym	Ndh-B1c {936}
Varieties	Fruskogorka.

Ndh-A1d

Synonym	Ndh-A1b {1037}
Varieties	Hope, Timgalen.

NDH-B1

Synonym	Ndh-A1 {513}
Chromosome	4BS
Varieties	CS.

NDH-D1

Chromosome	4DS
Varieties	CS.

NDH-E1

Chromosome	4E
ad	CS/ E. elongata .

NDH-H1

Synonym	Nadhd-1 {147}
Chromosome	4H
Chromosome	4HS
ad	CS/Betzes.

NDH-H[ch] 1

Chromosome	4H
ad	CS/ H. chilense .

NDH-V1

Chromosome	4V
ad	CS/ D. villosum .

NDH-R1

Chromosome	4RS
Chromosome	4R
ad	CS/Imperial, CS/King II {813, 362}; CS/Dakold {362}. NDH-S[1] 1 {813}. 4S[l] {813}.
ad	CS/ Ae. longissima .

NDH-U1

Note	A {362}.
ad	CS/ Ae. umbellulata .

MAL-E1

Chromosome	3E
ad	CS/ E. elongata .

MAL-H1

Chromosome	3H
ad	CS/Betzes.

MAL-R1

Chromosome	3R
ad	CS/Imperial.

GPT-A1

Chromosome	1AS
Varieties	CS.

GPT-B1

Chromosome

1BS

GPT-D1

Chromosome

1DS

GPT-E1

Chromosome	1ES
ad	CS/ E. elongata 1E.

GPT-H1

Chromosome	1H
dv	H. vulgare cv. Betzes.

CAT-B1

Synonym	Cat-A1 {1466}
Chromosome	4BL
Varieties	CS.

A catalase locus, designated CAT2 , was mapped 6 cM proximal to ACO-D2 in an Ae. tauschii F2 population derived from VIR-1954/VIR-1345 cross {10046}. This locus may be orthologous to CAT-B1 {10046}.

b-GLS

Note	2A[m] L {282}.
dv	DV92.

b-Glsb

Note	Null allele.
dv	G3116.

SBEI2

Chromosome	7BL
Varieties	CS {9937}.

SBEII

Suppression of SBEIIb expression alone had no effect on amylose contents; however, suppression of both SBEIIa and SBEIIb expression resulted in wheat starch containing >70% amylose {10534}. Combined loss-of-function mutations in SbeIIa-A, SbeIIa-B, SbeIIb-A , and SbeIIb-B (PI 670160) increased amylose content by 66% and resistant starch by 753% relative to the control in tetraploid wheat cv. Kronos {11125}. Combination of these four mutations with mutations of SbeIIa-D in hexaploid wheat (PI 670160) increased amylose content by 63% and resistant starch by 1,057% in field experiments relative to the control {11126}

PPO-B1

Varieties

Chinese Spring {10658}.

Ppo-B1a

Varieties	Chinese Spring {10658}.
c	GQ303713 {10658}.

PPO-D1

Marker associations

Detected with primers PPO16 and PPO29. Xwmc41-2D – 2.0 cM – PPO-D1 {10386}; Xcfd62-2D – 0.2 cM – PPO-D2 – 0.4 cM – Xcfd168-2D –

Ppo-D1a

Varieties	Chinese Spring {10386}; Louise {10931}; Zhonghou 9507 {10504, 10386}; others {10504, 10386}.
c	EF070149 {10386}. Wheats with this allele tend to have lower PPO activity {10386}.

Ppo-D1b

Note	EF070150 {10386}.
Varieties	CA 9632 {10386}; CA 9719 {10386}; Nongda 183 {10504}; others {10504, 10386}.
c	EF070150 {10386}. Wheats with this allele tend to have higher PPO activity {10386}.

Ppo-D1c

dv	Ae. tauschii Ae38 {10657}.
c	EU371656 {10657}.

Ppo-D1d

dv	Ae. tauschii Y59 {10657}.
c	EU371657 {10657}.

Ppo-D1e

Synonym	Ppo-D1null {10504}, Ppo-D1c {10656}
Varieties	Gaiyuerui {10504}; Zm2851 {10504}; XM2855 {10504}; 9114 {10504}. Wheats with this allele tend to have lower PPO activity {10504}.

PDI-1 PDI-A1

Chromosome	4AL
Varieties	CS {10422}.

PDI-B1

Chromosome	4DS
Varieties	CS {10422}.

PDI-D1

Chromosome	4BS
Varieties	CS {10422}. The genes for PDI and their promoters were sequenced in {10423}. A related sequence on 1BS was shown to be a partial, non-expressed

ISO-1

Synonym	ISA-1 {10295}
dv	Ae. tauschii {10295}.

PGIP-A1

Note	Tapgip3 , AM180658 {10608}.
dv	T. monococcum PI 538722 {10608}. Not expressed in T. urartu PI 428315 (AM884191) {10608} or in polyploid wheat because of inactivation by an inserted copia

PGIP-B1

Synonym	Tapgip1 {10610}
Chromosome	7BS
Marker associations	XS13M50-7B - 5 cM - PGIP-B1 - 11.7 cM - Xmgb105s-7B {10608}.

Pgip-B1a

Note	Tapgip1a {10608}.
Type varieties	Messapia {10608}.

Pgip-B1b

Note	Tapgip1b , AM884195 {10608}.
Type varieties	T. turgidum ssp. dicoccoides MG4343 {10608}. This non-expressed allele produces a large amplicon in southern blots using the Pgip sequence as probe, due to an

PGIP-D1

Synonym	Tapgip2 {10610}
Chromosome	7DS
Type varieties	Langdon 7D(7A) {10610}; Langdon 7D(7B) {10610}.

Pgip1

Chromosome	7BS
Varieties	CS ditelo 7BL {10390}.
Varieties (alt.)	Chinese Spring Pgip2 {10390}.
Type varieties	Langdon {10390}.

Pgip2

Chromosome	7DS
Varieties	CS ditelo 7DL {10390}.
Varieties (alt.)	Chinese Spring Pgip1 {10390}.

F3H-B1

Chromosome	2BL
Varieties	CS {10823}.
Marker associations	F3H-B1/Xgwm1067-2B – 11.4 cM – Xgwm1070-2B {10823}. ALLELIC VARIATION

F3H-D1

Chromosome	2DL
Varieties	CS {10823}.
Marker associations	Xgwm877-2D – 1.8 cM – F3HD1/Xgwm1264-2D – 22.7 cM – Xgwm301-2D {10823}. ALLELIC VARIATION

F3H-2 F3H-B2

Chromosome	2BL
Varieties	CS {10823}.
Marker associations	Xgwm1070-2B – 30.1 cM – F3H-B2 {10823}; Located in the terminal region near Xgwm1027-2B {10823}. ALLELIC VARIATION

ZDS-A1

Chromosome	2A
Type varieties	Langdon {10905}.

ZDS-B1

Chromosome	2B
Type varieties	Langdon {10905}.

ZDS-D1

Chromosome	2DL
Varieties	CS {10906}.

Zds-D1a

Synonym	TaZDS-D1a {10906}
Varieties	CA9632 {10906}; Many Chinese wheats and 80 CIMMYT lines {10906}.

Zds-D1b

Synonym	TaZDS-D1b {10906}
Varieties	Ning 99415-8 {10906}; Zhengzhou 9023 {10906}; Zhongyou 9507 {10906}; Zhoumai 13 {10906}.

Cv. Zhongyou 9507 has lower yellow flour pigment content, preferred for Chinese steamed bread and dry Chinese noodles. A QTL in the Zds-D1a region explained 18.4% of the variation in yellow pigment content in Zhongyou 9507/CA 9632 {10906}.

HYD-A1

Chromosome	2AL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

HYD-B1

Chromosome	2BL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

HYD-D1

Chromosome	2DL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

HYD-2 HYD-A2

Chromosome	5AL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

HYD-B2

Chromosome	4BL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

HYD-D2

Chromosome	4DL
Type varieties	Kronos {10913}.
Varieties	UC1041 {10913}.

eLCY-A1a

Varieties

Chinese Spring (11713{; WAWHT2074 {11713}. GenBank EU649785.

eLCY-A1b

Varieties

Ajana {11713}. GenBank JX288762. Alleles a and b were distinguished by a CAPS marker based on a SNP at position 2,028 bp. This difference

e-LCY-B1 [{10654}]. E-LCY3B {10654}. 3B {10654}. GenBank EU649786. e-LCY-D1 [{10654}]. E-LCY3A {10654}. 3D {10654}. GenBank EU649787.

AMP-2

AMP-A2

Chromosome	4AL
Varieties	CS.

Amp-A2a

Varieties

CS.

Amp-A2b

Varieties

T. spelta IPSR 1220017.

AMP-B2

Chromosome	4BS
Varieties	CS

Amp-B2a

Varieties

CS.

Amp-B2b

Varieties

Timstein.

Amp-B2c

Varieties

Hope.

AMP-D2

Chromosome	4DS
Varieties	CS.

Amp-D2a

Varieties

CS.

Amp-D2b

Varieties

Sears' Synthetic IPSR 1190903.

Amp-D2c

Varieties	Bersee. AMP-Ag[i] 2 {703}. 4Ag[i ] {703}.
ad	Vilmorin27/ Th. intermedium .

AMP-E2

Chromosome	4E
ad	CS/ E. elongata .

AMP-H2

Chromosome	4H
ad	CS/Betzes. AMP-H[ch] 2 {703}. 4H[ch ] {703}.
ad	CS/ H. chilense .

AMP-J2

Chromosome	4J
ad	CS/ Th. junceum . AMP-M[v] 2 {235}. 4M[v ] {235}.
su	H-93-33 {235}.

AMP-R2

Note	4RS {702}, {93}.
Chromosome	4R
ad	CS/Imperial. AMP-S[l] 2 {703}. 4S[l] L {703}.
ad	CS/ Ae. sharonensis {180}.
tr	4DS.4DL-4S[l] L {660}.

AMP-V2

Chromosome	4V
ad	CS/ D. villosum .

AMP-3

AMP-A3

Chromosome	7AS
Varieties	CS.

Amp-A3a

Varieties

CS.

Amp-A3b

Varieties

Timstein.

AMP-H3

Chromosome	7H
ad	CS/Betzes.

a-AMY-1

a-AMY-A1

Synonym

Amy6A {1082}

6AL {412}, {1082}.

a-Amy1a

Synonym	a-Amy-B1a
Varieties	CS.

a-Amy1b

Synonym	a-Amy-B1b
Varieties	CS.

a-AMY-A1

a-Amy-A1a

Synonym	Amy 6A1 {1084}
Varieties	CS. a-Amy-A1b[5] {7}.
Varieties	Bezostaya 1; Kavkaz. a-Amy-A1c[5] . [ Amy 6A1[m] {1084}].
Varieties	Aka.

a-AMY-B1

a-Amy-B1a

Synonym	Amy 4 {1084}, Amy 6B2[o] {1084}, Amy 6B1 {1084}
Varieties	CS {7}; Rare.

a-Amy-B1b

Synonym	Amy 4[m] {1084}, Amy 6B1[o ] {1084}, Amy 6B2 {1084}
Varieties	Mara {7}.

a-Amy-B1c

Synonym	Amy 6B1 {1084}, Amy 6B2 {1084}, Amy 4 {1084}
Varieties	Sava {7}; Rare.

a-Amy-B1d

Synonym	Amy 4[m ] {1084}, Amy6B2[o ] {1084}, Amy 6B1[o ] {1084}
Varieties	Sicco {7}; Rare.

a-Amy-B1e

Synonym	Amy 6B2[o ] {1084}, Amy 6B1[4' ] {1084}, Amy 4[m ] {1084}
Varieties	Cappelle-Desprez {7}.

a-Amy-B1f

Synonym	Amy 6B1[4] {1084}, Amy 6B2[o] {1084}, Amy4[m] {1084}
Varieties	Sappo {7}.

a-Amy-B1g

Synonym	Amy 4 {1084}, Amy 6B2[o] {1084}, Amy 6B1[4] {1084}
Varieties	Cheyenne {7}.

a-Amy-B1h

Synonym	Amy 6B2[o] {1084}, Amy 6B1[o] {1084}, Amy 4 {1084}
Varieties	T. macha Line 1 {7}; Rare.

a-AMY-D1

Synonym	Amy6D {1082}
Chromosome	6DL
Varieties	CS.

a-Amy-D1a

Synonym	Amy6D1 {1084}, Amy 6D2 {1084}
Varieties	CS.

a-Amy-D1b

Synonym	Amy 6D2 {1084}, Amy6D1 {1084}
Varieties	Prelude {1082}; Cappelle-Desprez {7}.

a-Amy-D1c

Synonym	Amy6D1[m] {1084}, Amy 6D2 {1084}
Varieties	T. spelta var. duhamelianum .

a-AMY-Ag[i] 1 {374}. 6Ag[i] {374}.

a-AMY-E1

Chromosome	6E
ad	CS/ E. elongata .

a-AMY-H1

Synonym	a-Amy1 {146}
Chromosome	6H
ad	CS/Betzes.

a-AMY-R1

Chromosome

6RL

su,ad: CS/Imperial; CS/King II; Holdfast/King II. a-AMY-R[m] 1 {13}. 6R[m] L {13}.

a-AMY-S1

Chromosome	6SS
Varieties	Wembley derivative 31.
al	Ae. speltoides .

Two types of nomenclature were assigned to the genes encoding the a-AMY-1 isozymes. In one, allelic states were defined for individual isozymes {1084} whereas in the other, several isozymes were considered the products of compound loci {7, 412}. This listing shows the 'alleles' described in {1084} which are assumed in {7} to be synonymous with the a- Amy-B1a through a-Amy-B1h nomenclature. Amy 4 and Amy 4[1] are unmapped alternatives {1084} which appear to be identical to zymogram bands [bands 9 and 9b {7}] forming part of the a-AMY-B1 phenotype. Amy 6B1 [with forms Amy 6B1[o] , and Amy 6B1[4'] , considered to be mutually exclusive {1084}] and Amy 6B2 [with forms Amy 62 and Amy 6B2[o] {1084}] describe further aspects of a-AMY-B1 {7}. See a-Amy1 below for further consideration of Amy 6B2 {1084}.

a-AMY-2

a-AMY-A2

Synonym	Amy7A {1082}
Chromosome	7AL

a-AMY-B2

Synonym

Amy7B {1082}

7BL {412}, {1082}.

a-Amy-B2a

Synonym	Amy 7B 1 {1084}, Amy 7B2 {1084}
Varieties	CS.

a-Amy-B2b

Synonym	Amy 7B1 {1084}, Amy 7B2[m] {1084}
Varieties	Hope. The alternative states of Amy 7B2 , namely, Amy 7B2 and Amy 7B2[m] {1084}, are identical to the variation in band 2 {412}. The

a-AMY-D2

Synonym	Amy7D {1082}
Chromosome	7DL
Varieties	CS.

a-Amy-D2a

Synonym	Amy 7D1 {1084}
Varieties	CS.

a-Amy-D2b

Synonym	Amy 7D1[o] {1084}
Varieties	Largo {7}; Sears' Synthetic {7}; VPM1 {417}. It was estimated {902} that there are two a-Amy-1 genes in chromosome 6A and five or

a-AMY-Ag[i] 2 {374}. 7Ag[i] {374}.

a-AMY-E2

Chromosome	7EL
ad	CS/ E. elongata .

a-AMY-H2

Synonym	a-Amy2 {146}
Chromosome	7HL
ad	CS/Betzes. a-AMY-H[ch] 2 {1015}. 7H[ch] beta {1015}. su,ad: CS/ H. chilense .

a-AMY-R2

Chromosome

7RL

su,ad: CS/Imperial; CS/King II; Holdfast/King II. a-AMY-S[b] 2 {13}. 7S[b] {13}.

a-AMY-U2

Chromosome	7U
ad	CS/ Ae. umbellulata . Three other a-Amy2 loci, namely, Amy 6B2, Amy 6D2, and Amy 7B2 , were reported {1084}. No variation was observed for

a-Amy1

Synonym	Amy 6B2 {1084}, Amy-B2 {1083}
Chromosome	6BL
Varieties	CS.

a-Amy1a

Synonym	a-Amy-B1a
Varieties	CS.

a-Amy1b

Synonym	a-Amy-B1b
Varieties	CS.

b-AMY-1

b-Amy-A1a

Synonym	b-Amy-A2a {8}, b-B1a {936}
Varieties	CS.

b-Amy-A1b

Synonym	b-B1b {936}, b-Amy-A2b {8}
Varieties	Koga II..

b-Amy-A1c

Synonym	b-B1c {936}, b-Amy-A2c {8}
Varieties	T. macha IPSR 1240005.

b-Amy-A1d

Synonym	b-Amy-A2d {8}, b-B1d {936}
Varieties	Holdfast.

b-Amy-A1e

Note	_
Synonym	b-Amy-A2e {8}, b-B1e_ {936}
Varieties	Bezostaya I.

b-AMY-B1

Synonym	b-Amy-A1 {8}
Chromosome	4BL
Varieties	CS.

b-Amy-B1a

Synonym	b-Amy-A1a {1330}}, {8}
Varieties	CS.

b-Amy-B1b

Synonym	b-Amy-A1b {1330}, {8}
Varieties	Sears' Synthetic IPSR 1190903.

b-Amy-B1c

Synonym	b-Amy-A1c {1330}, b-Amy-A1b {8}
Varieties	Ciano 67.

Amy-B1d

Synonym	b-Amy-A1c {1330}, {400}
Varieties	Manella.

b-AMY-D1

Chromosome	4DL
Varieties	CS.

b-Amy-D1a

Varieties

CS.

b-Amy-D1b

Varieties

Bersee.

b-Amy-D1c

Varieties

Sears' Synthetic. Rare. Previously listed alleles b-Amy-D1d and -D1e were found to be b-Amy-B1 alleles {400}. Two b-Amy-D[t] 1 alleles were predominant in 60 accessions

b-AMY-Ag[i] 1 {168}, {13}. 4Ag[i ] {168}.

b-AMY-C1

Note	B {1278}.
ad	Aestivum cv. Alcedo / Ae. caudata line B.

b-AMY-E[b] 1

Note	5E[b] L {661}.
tr	5AS.5E[b] L.

b-AMY-H1

Chromosome	4H
ad	CS/Betzes.

b-AMY-H[ch] 1

Chromosome	4H
ad	CS/ H. chilense .

b-AMY-R1

Synonym	b-AmyR1 {43}, b-Amy-R2 {13}
Chromosome	5R
Chromosome	5RL
ad	FEC 28/Petkus {43}; Holdfast/King II {43}, {1280}.
tr	CS/Imperial 5BL-5RL {43}.

b-AMY-S[l] 1

Note	D {13}.
Chromosome	4S
ad	CS/ Ae. sharonensis D {13}.
su	CS/ Ae. sharonensis .
ad	CS/ Ae. longissima .

b-AMY-U1

Synonym	b-Amy-U2 {13}
Chromosome	5U
su	CS/ Ae. umbellulata .

A second set of loci with homology to b-Amy-1 genes was identified in 2AS, 2BS and 2DS and designated the Xb-Amy-2 [ b-Amy-2 {1331}] set. Evidence for these genes derived from crosshybridization with a b-AMY-H1 cDNA probe {1331}. Further members of the same set were identified in 2H {732}, and 2R and 2U {1331}.

EP-2

EP-B2

Chromosome

6BS

An Ep locus was located in 4RS in King II rye {1280}, using Holdfast/King II addition lines and in 4R in Imperial {266} using Chinese Spring/Imperial addition lines.

EST-1

EST-A1

Synonym	EstA {61}
Chromosome	3AS
Varieties	CS.

EST-B1

Synonym	EstB {61}
Chromosome	3BS
Chromosome	3B
Varieties	CS.

EST-D1

Synonym	EstD {61}
Chromosome	3D
Chromosome	3DS
Varieties	CS.

Each of 208 hexaploid accessions carried the same Est-1 allele except accessions of T. compactum var. rubriceps , each of which carried an Est-B1 or Est-D1 electrophoretic mobility variant {585}.

EST-E1

Chromosome	3ES
ad	CS/ E. elongata .

EST-H1

Chromosome	3H
ad	CS/Betzes.

EST-R1

Synonym	EstR {61}
Chromosome	3R
ad	CS/Imperial {60}; Holdfast/King II {100}; Kharkov/Dakold {100}.

EST-S[1] 1

Chromosome	3S
ad	CS/ Ae. longissima .

EST-2

EST-A2

Synonym	Est-2A {585}
Chromosome	3A
Varieties	CS.

EST-B2

Synonym	Est-2B {585}
Chromosome	3BL
Varieties	CS. Among 208 hexaploid accessions, an apparent Est-B2 null allele occurred frequently in accessions of T. macha and T. sphaerococcum and occasionally in accessions of

EST-D2

Synonym	Est-2D {585}
Chromosome	3DL
Varieties	CS.

EST-3

EST-B3

Synonym	Est-3B {585}
Chromosome	7BS
Varieties	CS.

EST-D3

Synonym	Est-3D {585}
Chromosome	7DS
Varieties	CS.

EST-H3

Chromosome	7H
ad	CS/Betzes. One accession carrying an apparent Est-B3 null allele and one carrying an apparent Est-D3 null allele were found among 208 hexaploid accessions {585}.

A 7AS locus encodes three esterase isozymes in immature grains {009}.

EST-4

EST-A4

Synonym	Est-4A {585}
Chromosome	6AL
Varieties	CS.

EST-B4

Synonym	Est-4B {585}
Chromosome	6BL
Varieties	CS.

EST-D4

Synonym	Est-4D {585}
Chromosome	6DL
Varieties	CS.

Probable Est-A4 and Est-D4 null alleles were detected in several accessions of T. compactum var. rubriceps {585}; otherwise, no Est-4 variant occurred among 208 hexaploid accessions {585}. An esterase gene was located in chromosome L7 (= 6Ag[i] ) of Th. intermedium {361}.

EST-5

EST-5 consists of 20 or more monomeric, grain-specific isozymes that electrofocus between pH 5.6 and 7.0.

Est-A5a

Chromosome	3AL
Varieties	CS.

Est-A5b

Varieties

Kalyansona{9}; T. compactum AUS12084{756}.

EST-B5

Chromosome	3BL
Varieties	CS.

Est-B5a

Varieties

CS.

Est-B5b

Varieties

Big Club.

Est-B5c

Varieties

Timstein.

Est-B5d

Varieties

Sears' Synthetic.

EST-D5

Chromosome	3DL
Varieties	CS.

Est-D5a

Varieties

Est-D5b

Varieties

T. macha .

Est-D5c

Varieties

Hobbit 'S'.

Est-D5d

Varieties

T. macha Line 1.

Est-D5e

Varieties

T. macha WJR 38548. Sixty Ae. tauschii lines revealed six Est-D[t] 5 alleles {1578}.

Encoding of the endosperm esterases of hexaploid wheat by 12-15 genes in five compound loci located in 3AL, 3BL, 3DL, 3AS and 3DS was postulated in {1204}. Three and six alleles at Est-D[t] 5 (in Ae. tauschii ) were reported in {756} and {1578}, respectively. In S. cereale , in addition to EST-R1 , genes encoding leaf esterases were located in three chromosomes {1561}. These included a gene designated EST8 in 6R in cvs. Imperial and King II, a gene designated EST2 and two genes, designated EST6 and EST7 , which are part of a separate compound locus {1560}, in 5RL in Imperial, and a gene designaged EST10 in 4R of King II and 4RL of Imperial. In Hordeum vulgare , genes encoding leaf esterases were in 3H {1071; see also, 520,580} and 7H {520}.

EST-Ag[i] 5

Note	3Ag[i] {374}.
ad	Vilmorin 27/ Th. intermedium .

EST-H5

Chromosome	3H
ad	CS/Betzes.

EST-H[ch] 5

Chromosome	3H
ad	CS/ H. chilense .

EST-R5

Synonym

EstA {737}

6R {43}, {1280}.

EST-R[m] 5

Synonym	EstB {737}
Chromosome	6R
ad	CS/ S. montanum .

EST-S[b] 5

Chromosome

su,ad: CS/ Ae. bicornis .

EST-S[l] 5

Chromosome	3S
ad	CS/ Ae. longissima .

EST-6 EST-6 is a dimeric enzyme that electrofocuses around pH 7.6 and is specific to endosperm.

EST-A6

Chromosome	2AS
Varieties	CS.

Est-A6a

Varieties

CS.

Est-A6b

Varieties

Ceska Previvka.

EST-B6

Chromosome	2BS
Varieties	CS.

Est-B6a

Varieties

CS.

Est-B6b

Varieties

Hope.

EST-D6

Chromosome	2DS
Varieties	CS.

Est-D6a

Varieties

CS.

Est-D6b

Varieties

Sears' Synthetic IPSR 1190903.

EST-M6

Chromosome	2MS
su	CS/ Ae. comosa .

EST-R6

Chromosome	2RS
al	DS2 x RxL10 rye popn.

A group of leaf esterase isozymes controlled by the long arms of the homoeologous group 3 chromosomes were reported {919}. The relationship of these esterases to EST-2 and to the leaf esterase designed EST-6 reported in {629} was not determined.

EST-7

EST-A7

Chromosome	2AL
Varieties	CS.

EST-B7

Chromosome	2BL
Varieties	CS.

EST-D7

Chromosome	2DL
Varieties	CS.

Est-D7a

Varieties

CS.

Est-D7b

Varieties

Synthetic {IPSR 1190903}.

EST-E7

Note	2[E] {812}.
ad	CS/ E. elongata .

EST-H7

Chromosome	2HL
ad	CS/Betzes.

EST-R7

Chromosome	2RL
ad	CS/Imperial.
su	Holdfast/KingII.

EST-R[m] 7

Note	2R[m] alpha {812}.
ad	CS/ S. montanum .

EST-U7

Chromosome	2U
ad	CS/ Ae. umbellulata .

EST-V7

Chromosome	2V
ad	CS/ D. villosum .

EST-8 EST-8 consists of about 10 isozymes that electrofocus between pH 4.5 and 6.5 and are expressed only in vegetative tissues. EST-8 is likely to be the enzyme previously described in {919} and {629}. EST-A8 {629}, {814}. [ Est-A6 {629}]. 3AL {629}.

EST-A9

Chromosome	3AS
Varieties	CS.

EST-B9

Chromosome	3BS
Varieties	CS.

EST-D9

Chromosome

3DS

LPX-1

LPX-A1

Synonym	Lpx-B1 {516}
Chromosome	4AL
Varieties	CS {516}.
Marker associations	Xksu919(Lpx-1)-4A {0091}.

LPX-B1

Synonym	Lpx-A1 {516}
Chromosome	4BS
Varieties	CS {516}.
Marker associations	Xcn110(Lpx-1)-4B {0367, 0269}.

Lpx-B1a

Synonym	Lpx-A1a {936}
Varieties	CS.

Lpx-B1b

Synonym	Lpx-A1b {936}
Varieties	Bosanka {1533}.

LPX-B1.1

Chromosome	4BS
Marker associations	Xksm62-4B – 8 cM – LpxB1.1 – 13 cM – Xwmc617b-4B {10303}.

Lpx-B1.1a

Type varieties

UC1113 {10303}.

Lpx-B1.1b

Type varieties

Kofa, deletion {10303}.

LPX-B1.2

Chromosome	4B
Varieties	CS.

LPX-D1

Chromosome	4DS
Varieties	CS.

LPX-E1

Chromosome	4ES
ad	CS/ E. elongata .

LPX-H1

Chromosome	4H
ad	CS/Betzes.

LPX-2

LPX-A2

Note	5AL {516},{10303}.
Varieties	CS.
Marker associations	Xksu919(Lpx-2)-5A {91}.

LPX-B2

Note	5BL {516},{10303}.
Varieties	CS.
Marker associations	Xksu919(Lpx-2)-5B {91}; Xcn111(Lpx-2)-5B {269}.

LPX-D2

Chromosome	5DL
Varieties	CS.

LPX-E2

Chromosome	5EL
ad	CS/ E. elongata .

LPX-H2

Chromosome	5H
ad	CS/Betzes.

LPX-S[s] 2

Chromosome	5S
ad	CS/ Ae. searsii .

LPX-V2

Chromosome	5V
ad	CS/ D. villosum .

LPX-3 LPX-A3

Chromosome	4AL
Type varieties	UC1113 (GenBank DQ474244) and Kofa (GenBank DQ474242) {10303}.
Marker associations	Xwmc617a-4A – 10 cM – Lpx-A3 – 15 cM – Xgwm192b-4A {10303}.

LPX-B3

Chromosome	4B
Type varieties	UC1113 and Kofa (GenBank DQ474243) {10303}.

AADH-1

AADH-A1

Synonym	Adh-A2 {584}
Chromosome	5AL
Varieties	CS.
Marker associations	XksuG44-5A (proximal) - 6.9 cM - AADH-A1 - 24.7 cM - Xcdo412-5 (distal) {9959}.

Aadh-A1a

Varieties

CS; 133 other accessions {584}.

Aadh-A1b

Varieties

T. spelta ; K-24696; other accessions {584}.

AADH-B1

Synonym	Adh-B2 {584}
Chromosome	5BL
Varieties	CS.

Aadh-B1a

Varieties

CS {1533}.

Aadh-B1b

Varieties

Drina {1533}.

AADH-D1

Synonym	Adh-D2 {584}
Chromosome	5DL
Varieties	CS.

AADH-C1

Note	C {1278}.
ad	Alcedo/ Ae. caudata line C.

AADH-E1

Synonym	Adh-E2 {518}
Chromosome	5EL
ad	CS/ E. elongata .

AADH-R1

Chromosome	5RL
ad	Holdfast/King II.

AADH-2

AADH-A2

Synonym	Adh-A3 {508}
Chromosome	6A

6AL {513}, {587}.

AADH-B2

Synonym	Adh-B3 {508}
Chromosome	6B
Chromosome	6BL
Varieties	CS {513}; Carola {1279}.

AADH-D2

Synonym	Adh-D3 {508}
Chromosome	6DL
Chromosome	6D
Varieties	CS {513}; Carola {1279}.

AADH-Ag[e] 2

6Ag[e ] {1575}. ad,su: Rescue/ Th. elongatum .

AADH-E2

Synonym

Adh-E3 {518}

6EBeta {518}.

AADH-R2

Chromosome	6RL
ad	Holdfast/King II.

AADH-V2

Chromosome	6V
ad	CS/ D. villosum .

The AADH-1 and AADH-2 loci were designated with the synonyms Adh-2 and Adh-3 , respectively, in some publications in addition to {508}, {518}, {584}. These include: {510}, {509}, {511}, {519}, {517}, {587}, 1066, 1139}.

NDH-2

NDH-A2

Chromosome	7A
Varieties	Hope.

NDH-D2

Chromosome	7DS
Varieties	CS.

NDH-R2

Chromosome	7RS
ad	CS/Imperial, CS/King II, Holdfast/King II (7R).

NDH-3 NDH-A3

Chromosome

3AL

NDH-B3

Chromosome	3BL
Varieties	CS.

Ndh-B3a

Varieties

CS.

Ndh-B3b

Varieties

Carmen.

NDH-D3

Chromosome	3DL
Varieties	CS.

A NDH locus, designated NADHD2 , was mapped 27 cM from Est-D10 in an Ae. taushii F2 population derived from VIR-1954/VIR-1345 {10046}. This locus may be homologous to NDH-D3 .

NDH-H3

Chromosome	3HL
ad	CS/Betzes.

NDH-R3

Chromosome	6RL
ad	Holdfast/King II, CS/Imperial (6R), CS/King II (6R).

NDH-S[l] 3

Note	3S[l] L {813}.
ad	CS/ Ae. longissima ; CS/ Ae. sharonesis (3S[l] ).

Based on the correspondence of the electrophoretic patterns, isoelectric points (pIs) and chromosomal location, it was proposed that NDH-3 (NADH dehydrogenase), DIA1 (diaphorase) and MNR1 (menadione reductase) represent the same locus {0356}.

NDH-4 NDH-A4

Chromosome	3AS
Varieties	CS.

NDH-B4

Chromosome	3BS
Varieties	CS.

NDH-E4

Chromosome	3ES
ad	CS/ E. elongata .

NDH-H4

Chromosome	3HS
ad	CS/Betzes.

NDH-R4

Chromosome	3RS
ad	CS/King II, CS/Imperial (3R).

DIP-B1

Synonym	Pept-B1 {1533}
Chromosome	6BL
Varieties	CS.

Dip-B1a

Varieties

CS.

Dip-B1b

Varieties

Cappelle-Desprez.

DIP-D1

Chromosome	6DL
Varieties	CS.

DIP-H1

Synonym

Pept-1 {147}, Dip 1 {145}

6H {145}, {147}, {700}.

DIP-J1

Chromosome	6J
ad	CS/ Th. junceum .

DIP-V1

Chromosome	6V
ad	CS/ D. villosum .

AHASL 1

AHASL-A1

Synonym	Imi3 {10099}
Chromosome	6AL
Varieties (alt.)	CDC Teal IMI 15A Imi3 {10099}.
dv	T. monococcum mutant EM2 (mutant of susceptible line TM23 {10102}.

AHASL-B1

Synonym	Imi2 {10099}
Chromosome	6BL
Varieties	CDC Teal IMI 11A = PTA3953 {10099}.

AHASL-D1

Synonym	Imi1 {10099}
Chromosome	6DL
Varieties	BW755 = Grandin*3/Fidel-Fs-4 {10099}.

PSY-1

PSY1-A1

Chromosome	7AL
Type varieties	Kofa {10230}.
Marker associations	Xwmc809-7A – 5.8 cM – Yp7A {10501}.

Psy1-A1a

Type varieties	Kofa {10230}.
Marker associations	Xwmc809-7A - 5.8 cM - Yp7A {10501}.

Psy1-A1b

Varieties	Chinese common wheats with low yellow pigment content {10501}; PH82-2 {10501}; Shaan 9314 {10501}; Xinong 336 {10501}.
c	GenBank EF600064 {10501}. 37-bp insertion in intron 2 (231 bp fragment for marker Yp7A ) {10501}. 676-bp insertion in intron 4 {10530}.

Psy1-A1c

Varieties	M564 {10650}.
c	GenBank EU650391 {10650}; No 37-bp insertion in intron 2 and no 676-bp insertion in intron 4 {10530}; High yellow pigment cultivars: Aroona (PI

Psy1-A1d

Type varieties	Langdon {10651}; T. dicoccum DM28 {10652}.
c	GenBank EU263018 {10651}; FJ393515 {10652}.

Psy1-A1e

Varieties	Sunco {10654}.
Type varieties	DR8 {10651}.
c	EU649791 {10654}; EU263019 {10651}.

Psy1-A1f

dv	T. urartu PI 428326 {10652}.
c	FJ393516 {10652}.

Psy1-A1g

dv	T. urartu UR1 {10652}.
c	FJ393517{10652}.

Psy1-A1h

dv	T. boeoticum BO1 {10652}; T. monococcum MO5 {10652}.
c	FJ393518 {10652}; FJ393519 {10652}.

Psy1-A1i

dv	T. monococcum MO1 {10652}.
c	FJ393520 {10652}.

Psy1-A1j

dv	T. monococcum MO2 {10652}.
c	FJ393521 {10652}.

Psy1-A1k

Varieties	Spelt 167 {10652}.
Type varieties	T. dicoccoides DS3 {10652}; T. dicoccum DM37 {10652}.
c	FJ293527 {10652}; FJ293522 {10652}; FJ293523 {10652}.

Psy1-A1l

Type varieties	Kofa {10530, 10230}; Strongfield {10653}; T. dicoccoides DS6 {10652}.
c	EU096090 {10530, 10230}; FJ393524 {10652}.

Psy1-A1m

Type varieties	T. dicoccum DM26 {10652}.
c	FJ393525 {10652}.

Psy1-A1n

Varieties	Spelt SP9{10652}.
c	FJ393526 {10652}.

Psy1-A1o

Type varieties	Commander {10653}.
c	FJ234424 {10653}.

Psy1-A1p

Varieties	Tasman {10654}.
c	EU649792 {10654}.

Psy1-A1q

Varieties	Cranbrook {10654}.
c	EU649793 {10654}.

Psy1-A1r

Varieties	Halberd {10654}.
c	EU649794 {10654}.

Psy1-A1s

Varieties	Schomburgk {10654}.
c	EU649795 {10654}.

Psy1-A1t

Varieties	WAWHT2074 {10920}.
Marker associations	Xwgm344-7A - 3.9 cM - Psy1-A1t - 9.9 cM - Ccfa2257a-7A {10920}.
c	GenBank HM006895 {10920}.

PSY1-B1

Chromosome	7BL
Type varieties	Kofa{10230}.
Marker associations	Xcfa2040-7B – 12 cM – PSY1-B1 – 5 cM – Xgwm146-7B {10230}.

Psy1-B1a

Note	GenBank EU096093 {10530}.
Synonym	{10530}
Varieties	Chinese Spring {10654, 10650, 10530}; Spelt SP9 {10652}.
Type varieties	T. dicoccoides DS4 {10652}.
c	FJ393529 {10652}; FJ393528 {10652}; EU650392 {10650}; EU096094 {10530}; EU649789 {10654}.

Psy1-B1b

Varieties	Neixiang 188 {10650}.
c	EU650393 {10650}.

Psy1-B1c

Varieties	CA 9648 {10650}.
c	EU650394 {10650}.

Psy1-B1d

Varieties	Ning 98084 {10650}.
c	EU650395 {10650}.

Psy1-B1e

Varieties	M484 {10650}.
Type varieties	DR8 {10650}; T. dicoccum DM28 {10652}.
c	EU263021{10650}; FJ393541 {10652}.

Psy1-B1f

Type varieties	Langdon {10651}.
c	EU263020 {10651}.

Psy1-B1g

Type varieties	DR1 {10651}; T. dicoccoides DS6 {10652}.
c	EU650396 {10651}; FJ393530 {10652}.

Psy1-B1h

Type varieties	T. dicoccoides DS3 {10652}.
c	FJ393531 {10652}.

Psy1-B1i

Type varieties	T. dicoccoides DS8 {10652}.
c	FJ393532 {10652}.

Psy1-B1j

Type varieties	T. dicoccum DM26 {10652}.
c	FJ393533 {10652}.

Psy1-B1k

Type varieties	T. dicoccum DM33 {10652}.
c	FJ393534 {10652}.

Psy1-B1l

Type varieties	T. dicoccum DM37 {10652}.
c	FJ393535 {10652}.

Psy1-B1m

Varieties	Spelt 167 {10652}.
Type varieties	T. dicoccum DM47 {10652}.
c	FJ393540 {10652}; FJ393539 {10652}.

Psy1-B1n

Note	Previously designated Psy1-B1b {10656}.
Type varieties	Kofa.
c	EU096092 {10530}; DQ642439 {10230}.

Psy1-B1o

Note	Previously designated Psy1-B1a {10656}.
Type varieties	UC1113 {10530}; W9262-260D3 {10230}.
c	EU096093 {10530}; DQ642440 {10230}.

PSY1-D1

Chromosome

7DL

Psy1-D1a

Varieties	Chinese Spring {10652}.
c	EU650397 {10652}; EU649790 {10654}.

Psy1-D1b

dv	Ae. tauschii Ae34 {10652}.
c	FJ393542 {10652}.

Psy1-D1c

dv	Ae. tauschii Ae46 {10652}.
c	FJ393543 {10652}.

Psy1-D1d

dv	Ae. tauschii Y99 {10652}.
c	FJ393544 {10652}.

Psy1-D1e

Varieties	Spelt SP9 {10652}.
c	FJ393545 {10652}.

Psy1-D1f

Varieties	Spelt 217 {10652}.
c	FJ393546 {10652}.

Psy1-D1g

Varieties	Zhonliang 88375 {10652}.
c	FJ807498 {10652}.

Psy1-D1h

dv	Ae. tauschii Ae37 {10652}.
c	FJ807499 {10652}.

Psy1-D1i

dv	Ae. tauschii Ae38 {10652}.
c	FJ807500{10652}.

Psy1-D1j

dv	Ae. tauschii Ae42 {10652}.
c	FJ807501 {10652}.

Psy1-D1k

Varieties	Nongda 3291 {10655}.
c	FJ807495 {10655}.

Psy1-D1l

Varieties	E 86642 {10655}.
c	FJ807496 {10655}.

Psy1-D1m

Varieties	Ning 97-18 {10655}.
c	FJ807497 {10655}.

PSY1-E1

Synonym

Psy-E1

Psy1-E1a

Note	GenBank EU096095 {10530}.
Synonym	Psy-E1a {10530}
Varieties	Agatha (7EL translocation) {10530}.

Psy1-E1b

Synonym

Psy-E1b {10530}

Similar to EU096095, but with P to L substitution at amino acid 422 {10530}.

PSY1-S1

al	Ae. speltoides Ae48 {10652}.

Psy1-S1a

al	Ae. speltoides Ae48 {10652}.
c	FJ393536 {10652}.

Psy1-S1b

al	Ae. speltoides Ae49 {10652}.
c	FJ393537 {10652}.

Psy1-S1c

al	Ae. speltoides Y162 {10652}.
c	FJ393538 {10652}.

PSY-2 Homology with the same gene in rice ( Psy2 ) {10230}.

PSY2-A1

Chromosome	5A
Type varieties	Kofa {10230}.

PSY2-B1

Chromosome	5B
Type varieties	Kofa {10230}.
Marker associations	Xgwm191-5B – 17 cM – PSY-B2 {10230}.

PPO-2

PPO-A2

Synonym	PPO-A2 {10931}
Chromosome	2AL
Marker associations	Xcfa2058-2A – 0.4 cM – PPO-A2 – 0.4 cM – Xiwa174-2A – 8.3 cM – Xiwa7593-2A – 0.6 cM – PPO-A1 – 11.0 cM –

Ppo-A2a

Varieties	Alpowa {10930}.
c	GenBank HQ228148 {10930}.

Ppo-A2b

Varieties	Panawawa {10931}.
c	GenBank HQ228149 {10930}.

Ppo-A2c

Varieties	Louise {10931}.
c	JN632507 {10931}.

PPO-B2

Synonym	PPO-B2 {10930}
Chromosome	2B
Marker associations	Xiwa175/Xiwa4866-2B - 0.7 cM - PPO-B2 - 2.3 cM - Xiwa7593-2B {10931}.

Ppo-B2a

Varieties	Penawawa {10931}.
c	GenBank HQ228150 {10930}.

Ppo-B2b

Varieties	Alpowa {10930}.
c	GenBank HQ228151 {10930}.

Ppo-B2c

Varieties	Louise {1211}.
c	GenBank JN632508 {10930}.

PPO-D2

Synonym	PPO-D2 {10930}
Chromosome	2DL
Marker associations	Xcfd62-2D – 0.2 cM – PPO-D2 – 0.4 cM – Xcfd168-2D – 7.7 cM – Xgwm608-2A – 2.6 cM – PPO-D1 – 0.9 cM –

Ppo-D2a

Varieties	Louise {10931}.
c	GenBank HQ228152 {10931}.

Ppo-D2b

Varieties	Penawawa {10930}.
c	HQ228153 {10930}. Wheats with this allele tend to have lower PPO activity {10385, 10386}.

2.3. Endosperm storage proteins⌂ Home

Glu-A1a

Note	1 {1116}.
Varieties	Hope.

Glu-A1b

Note	2[* ] {1116}.
Varieties	Bezostaya 1.

Glu-A1c

Note	Null allele {1116}.
Varieties	CS.

Glu-A1d

Varieties

V74, Spain {1115}.

Glu-A1e

Varieties

132c, Poland {1115}.

Glu-A1f

Varieties

112-29, Sudan {1115}.

Glu-A1g

Varieties

Landrace 1600.

Glu-A1h

Synonym	GLU-A1-I {1527}
Type varieties	PI 94683, USSR, T. dicoccum .

Glu-A1i

Synonym	GLU-A1-II {1527}
Type varieties	CI 12213, India, T. dicoccum ; Lambro {1523}.

Glu-A1j

Synonym

GLU-A1-III {1527}

1' {125}.

Glu-A1k

Note	26 {478}.
Varieties	BT-2288 {478}.

Glu-A1l

Type varieties

Chinook, Canada.

Glu-A1m

Type varieties

Nugget Biotype 1, Canada, T. durum .

Glu-A1n

Synonym

Glu-A1m {959}, Glu A1-IV {1526}

1' {125}.

Glu-A1p

Note	3[* ] {1146}.
Varieties	David 1.

Glu-A1q

Synonym

Glu A1VI {125}

2[***] {125}.

Glu-A1r

Note	39+40 {1232}.
i	T. thaoudar IPSR 1020006/6[*] Sicco.

Glu-A1s

Note	41+42 {1231}.
i	T. thaoudar G3152/6[*] Sicco.

Glu-A1t

Note	21[*] {602}.
Varieties	W29323, W3879, W31169.

Glu-A1u

Note	2
Synonym	B] {02106}. v: Bankuti 1201. The allele designated Glu-A1u* and Glu-A1-1u in the appropriate list below encodes a high molecular weight glutenin subunit (denominated 2[*B]

[ Glu-A1-VII {03137}]. VII {03137}.

Glu-A1w

Note	2.1* {10327}
Varieties	KU-1094, KU-1026, KU-1086, Grado, KU-1139 {10327}.

Glu-A1x

Note	2' {10327}.
Varieties	TRI14165/91 {10327}. The alleles formerly designated t to x in {959} were renamed x to ab because allele t in {847} and alleles u, v

Glu-A1y

Synonym

2'' {10535}

**v:

Glu-A1ac

Synonym	Glu-A[u] 1-I {10806}
dv	PI 428319 {10806}.

Glu-A1ad

Synonym	Glu-A[u] 1-II {10806}
dv	PI 428232 {10806}.

Glu-A1ae

Synonym	Glu-A[u] 1-III {10806}
dv	PI 428240 {10806}.

Glu-A1af

Synonym	Glu-A[u] 1-IV {10806}
dv	PI 428335 {10806}.

Glu-A1ag

Synonym	Glu-A[u] 1-V {10806}
dv	PI 538741 {10806}.

Glu-A1ah

Synonym	Glu-A[u] 1-VI {10806}
dv	PI 428230 {10806}.

Glu-A1ai

Synonym	Glu-A[u] 1-VII {10806}
dv	PI 428253 {10806}.

Glu-A1aj

Synonym	Glu-A[u] 1-VIII {10806}
dv	PI 427328 {10806}.

Glu-A1ak

Synonym	Glu-A[u] 1-IX {10806}
dv	PI 428327 {10806}.

Glu-A1al

Synonym	Glu-A[u] 1-X {10806}
dv	PI 428256 {10806}.

Glu-A1am

Synonym	Glu-A[u] 1-XI {10806}
dv	PI 428224 {10806}.

Glu-A1an

Synonym	Glu-A[u] 1-XII {10806}
dv	PI 428228 {10806}.

Glu-A1ao

Synonym	Glu-A[u] 1-XIII {10806}
dv	PI 538724 {10806}.

Glu-A1ap

Synonym	Glu-A[u] 1-XIV {10806}
dv	TRI 6734 {10806}.

Glu-A1aq

Synonym	Glu-A[u] 1-XV {10806}
dv	TRI 11494 {10806}.

Glu-A1ar

Synonym	Glu-A[u] 1-XVI {10806}
dv	TRI 11495 {10806}.

Glu-A1as

Synonym	Glu-A[u] 1-XVII {10806}
dv	PI 428217 {10806}.

Glu-A1at

Synonym	Glu-A[u] 3-XVIII {10806}
dv	PI 428225 {10806}.

Glu-A1au

Synonym	Glu-A[u] 3-XIX {10806}
dv	PI 538733 {10806}.

Glu-A1av

Synonym	Glu-A[u] 3-XX {10806}
dv	PI 428196 {10806}.

Glu-A1aw

Synonym	Glu-A[u] 3-XXI {10806}
dv	PI 538724 {10806}.

Glu-A1ax

Synonym	Glu-A[u] 3-XXII {10806}
dv	PI 428191 {10806}.

Glu-A1ay

Synonym	Glu-A[u] 3-XXIII {10806}
dv	TRI 6734 {10806}.

Glu-A1az

Synonym	Glu-A[u] 3-XXIV {10806}
dv	TRI 11496 {10806}.

Glu-A1ba

Synonym

Glu-A1g {11106}

1.1 {11106}.

GLU-B1

Synonym

Glt-B1 {420}, Glu 1B {1415}, Glt-B3 {420}, Glt-B2 {420}

1BL {1125}, {107}, {780}.

Glu-B1a

Note	7 {1116}.
Varieties	Flinor.

Glu-B1b

Note	7+8 {1116}.
Varieties	CS. Subunit 8 of Glu-B1b (7+8) is more acidic in isoelectric focusing than subunit 8 of Glu-B1d (6+8) {555}. Variation in the mobility of subunits

Glu-B1c

Note	7+9 {1116}.
Varieties	Bezostaya 1.

Glu-B1d

Note	6+8 {1116}.
Varieties	Hope.
Type varieties	Kronos {11497}. Simultaneous and individual truncation mutations were found in Glu-B1x and Glu-B1y subunits in Kronos mutant lines {11497}. Germplasm was accessioned as PI 692251

Glu-B1e

Note	20 {1116}.20x+20y {03133}.
Varieties	Federation.

Glu-B1f

Note	13+16 {1116}.
Varieties	Lancota (rare). Primers were designed to distinguish subunit By8 from By8*, for distinguishing subunit By9-containing alleles from non-By9 alleles, and for diagnosing the presence of

Glu-B1g

Note	13+19 {1116}.
Varieties	NS 335 (rare).

Glu-B1h

Note	14+15 {1116}.
Varieties	Sappo (rare).

Glu-B1i

Note	17+18 {1116}.
Varieties	Gabo. Although alleles Glu-B1i encoding subunits 17+18, and Glu-B1bc encoding subunit 6+17, apparently share a common subunit (Ax17 and By17, respectively) it is not clear

Glu-B1j

Note	21 {1116}.21x+21y {3116}.
Varieties	Dunav (rare); Foison {03116}.

Glu-B1k

Note	22 {1116}.
Varieties	Serbian (rare).

Glu-B1l

Note	23+24 {778}.
Varieties	Spica D.

Glu-B1m

Synonym	GLU-B1-I {1527}
Type varieties	PI 94640, Iran, T. dicoccum .

Glu-B1n

Synonym	GLU-B1-II {1527}
Type varieties	PI 355505, Germany, T. dicoccum .

Glu-B1o

Synonym	GLU-B1-III {1527}
Type varieties	PI 352354, Ethiopia, T. dicoccum .

Glu-B1p

Synonym

GLU-B1-IV {1527}

23+18 {125}.

Glu-B1q

Synonym	GLU-B1-V {1527}
Type varieties	PI 94633, Morocco, T. dicoccum .

Glu-B1r

Synonym

GLU-B1-VI {1527}

19 {125}.

Glu-B1s

Note	7+11 {478}.
Varieties	BT-2288. Subunit 11 of Glu-B1s (7+11) was so numbered in {478} because its mobility is the same as one of the subunits encoded by a

Glu-B1t

Varieties

Supreza, Canada.

Glu-B1u

Note	7[*] +8 {1146}.
Varieties	Owens {1069}; Fiorello {1146}.

Glu-B1v

Varieties

Mondor.

Glu-B1w

Note	6[] +8[ ] {1146}.
Varieties	Dawbull {1069}; Sieve {1146}.

Possible low gene expression at Glu-B1 was noted for Glu-B1w , where subunits 6[*] +8[*] stain very faintly {1146}.

Glu-B1x

Synonym	Glu-B1-VII {1526}, Glu-B1t {959}
Type varieties	Canoco de Grao Escuro, Portugal, T. turgidum .

Glu-B1y

Synonym	Glu-B1-VIII {1526}, Glu-B1u {959}
Type varieties	Tremez Mollez, Portugal, T. durum .

Glu-B1z

Synonym

Glu-B1-IX {1524}, Glu-B1v {959}

7+15 {125}.

Glu-B1aa

Synonym	Glu-B1w {959}, Glu-B1-X {1524}
Type varieties	Quaduro, Italy, T. durum .

Glu-B1ab

Synonym	Glu-B1x {959}, Glu-B1-XI {1523}
Type varieties	Athena, Italy, T. durum .

Glu-B1ac

Synonym

Glu B1XIII {125}

6+16 {125}.

Glu-B1ad

Synonym

Glu B1XIV {125}

23+22 {125}.

Glu-B1ae

Note	18[* ] {1146}.
Varieties	David.

Glu-B1af

Note	26+27 {1146}.
Varieties	Cologna 1. One of the Glu-B1af subunits was numbered 26 in {1146}; 26 was previously used to number the subunit encoded by Glu-A1k {478}.

Glu-B1ag

Note	28+29 {1146}.
Varieties	Forlani. Subunit 28 of Glu-B1ag (28+29) {1146} is referred to as subunit 19[*] in {1068}.

Glu-B1ah

Note	Null allele {782}.
Varieties	Olympic mutant.

Glu-B1ai

Note	7' {714}.
Varieties	Adonis.

Glu-B1aj

Note	8 {759}.
Varieties	AUS 14444, Afghanistan.

Glu-B1ak

Note	7[] +8[] {899}.
Varieties	Norstar.

Glu-B1al

Note	7[OE] +7[OE] +8[*] {899}.
Varieties	Benkuti 1201; Glenlea {899}; Klein Universal II {10196}; Tezanos Pintos Precoz {10196}; Tobari 66 {10196}. Other genotypes are listed in {10196}. Many of the

Glu-B1am

Note	18 {1229}.
Varieties	Royo.

Glu-B1an

Note	6 {1229}.
Varieties	BG-2013.

Glu-B1ao

Note	7+16 {1229}.
Varieties	BG-3545.

Glu-B1ap

Note	30+31 {1229}.
Varieties	Marinar.

Glu-B1aq

Note	32+33 {1229}.
Varieties	BG-1943.

Glu-B1ar

Note	34+35 {1229}.
Varieties	Jeja Almendros.

Glu-B1as

Note	13 {1229}.
Varieties	PI 348435.

Glu-B1at

Note	13+18 {1229}.
Varieties	PI 348449.

Glu-B1au

Note	37 {1032}.
Varieties	Shedraya Polesja.

Glu-B1av

Synonym

Glu-B1r {3116}

7-18 {03116}.

Glu-B1aw

Synonym

Glu-B1s {03116}

6.8-20y {03116}.

Glu-B1ax

Synonym

Glu-B1-XV {03137}

XV {03137}.

Glu-B1ay

Synonym

Glu-B1-XVI {03137}

XVI {03137}.

Glu-B1az

Synonym

Glu-B1-XVII {03137}

XVII {03137}.

Glu-B1ba

Synonym

Glu-B1-XVIII {03122}

13[*] +16 {03122}.

Glu-B1bb

Synonym

Glu-B1-XLX {03122}

6+18' {03122}.

Glu-B1bc

Note	6+17 {03138}.
Varieties	ICDW 20975 {03138}.

Glu-B1bd

Note	20+8 {03140}.
Varieties	Abadja {03140}.

Glu-B1be

Type varieties

T. dicoccoides Israel-A {10186}.

Glu-B1bf

Type varieties

T. dicoccoides PI 481521 {10186}.

Glu-B1bg

Type varieties

T. dicoccoides PI 478742 {10186}.

Glu-B1bh

Note	13+22 {10327}.
Varieties	Grado {10327}; KU-1026 {10327}; KU-1086 {10327}; KU1094 {10327}; KU-1139 {10327}.

Glu-B1bi

Note	13+22.1 {10327}.
Varieties	KU-1135 {10327}.

Glu-B1bj

Note	14+15 {10327}.
Varieties	TRI11553/92 {10327}.

Glu-B1bk

Synonym

Glu-B1be {10327}

6.1+22.1 {10327}.

Glu-B1bl

Synonym

Glu-B1bf {10327}

6.1 {10327}.

Glu-B1bm

Synonym

Glu-B1bg {10327}

13*+19* {10327}.

Glu-B1bn

Note	7+19 {10425}.
Varieties	Triticales: Lasko, Dagno, Tewo, Vision, Dato {10425}.

Glu-B1bo

Note	7+26 {10425}.
Varieties	Triticales: Presto, Modus {10425}. The number 26 was also used to designate a subunit encoded by Glu-A1k and Glu-A1-1k .

Glu-B1bp

Note	7[**] +8 {10643}.
Varieties	XM1368-2 {10643}; XM1404-2 {10643}.

Glu-B1bq

7+8[**] {10643}.

Glu-B1br

Note	7.1+7.2+8[*] {10807}.
Varieties	H45 {10807}.

Glu-B1bs

Note	7.3+7[OE] +8[*] {10807}.
Varieties	VQ0437 {10807}.

Glu-B1bt

Note	17'+18' {10809}.
Type varieties	TGR-214 {10809}.

Glu-B1bu

Note	17'+18[* ] {10809}.
Type varieties	TGR-2246 {10809}.

Glu-B1bv

Note	13[*] +8[] {10809}.
Type varieties	TGR-003 {10809}.

Glu-B1bw

Note	8' {10809}.
Type varieties	TGR-244 {10809}.

Glu-B1bx

Note	7+17 {10810}.
Varieties	CWI-59797, T. aestivum var. ferrugineum {10810}.

Glu-B1by

Note	7b[*] +8 {10808}.
Varieties	Eshimashinriki {10808}.

Glu-B1bz

Note	7[OE ] {10808}.
Varieties	Attila {10808}; Darius {10808}; Cappelle-Desprez {10808}; Festin {10808}; Petrel {10808}.

Glu-B1ca

Note	6+8b[* ] {10808}.
Varieties	Appolo {10808}; Brimstone {10808}; Clement {10808}; Nidera Baguette 10 {10808}; Ruso {10808}; Pepital {10808}; Thesee {10808}.

Glu-B1cb

Note	7[OE] +8 {10808}.
Varieties	ACA 303 {10808}; Courtot {10808}; Demai 3 {10808}; Shinchunaga {10808}.

Glu-B1cc

Note	7[OE] +8a[* ] {10808}.
Varieties	Klein Jabal 1 {10808}; Pioneer {10808}; ProINTA {10808}; Redemon {10808}.

Glu-B1cd

Note	7[OE] +8b[* ] {10808}.
Varieties	ACA 601 {10808}.

Glu-B1ce

Note	7+8a[* ] {10808}.
Varieties	Jing 411 {10808}; Tasman {10808}. In a study including emmer wheats ( T. dicoccon ) {00115}, new subunits named 7[+] (in accessions MG5400/5 and

Glu-B1cf

Note	20+33 {11490}.
Type varieties	T. turgidum ssp. durum Mexican landrace accession 22 (CWI52215) {11490}.

Glu-B1cg

Note	13+16* {11490}.
Type varieties	T. turgidum ssp. durum Mexican landrace accession 19 (CWI52200) {11490}.

Glu-B1ch

Note	7+22 {11490}.
Varieties	T. aestivum ssp. aestivum cv. Wilbur (CW13735) {11490}.

Glu-B1ci

Note	7+22* {11490}.
Type varieties	T. turgidum ssp. durum Iranian landrace accession 51 (CWI57280) {11490

Glu-B1cj

Note	13+15 {11490}.
Type varieties	T. turgidum ssp. durum Iranian landrace accession 46 (CWI56913) {11490}.

Glu-B1ck

Note	15 {11491}.
Varieties	T. aestivum ssp. compactum PI 157920 {11491}.

Glu-B1cl

Note	14+8 {11491}.
Varieties	T. aestivum ssp. macha PI 272554, PI 278660, PI 290507 {11491}.

Glu-B1cm

Note	6+8* {11491}.
Varieties	T. aestivum ssp. macha PI 428177 {11491}.

Glu-B1cn

Note	17 {11491}.
Varieties	T. aestivum ssp. sphaerococcum CItr 4531, PI 272581, PI 282452 {11491}.

Glu-B1co

Note	20+22* {11493}.
Type varieties	T. turgidum ssp. durum Moroccan landraces MGB-2963, MGB-3152 {11493}.

Glu-B1cp

Note	20* {11493}.
Type varieties	T. turgidum ssp. durum North American cv. MGB-66023 {11493}. Glu-B1cq [{11492}]. 7+8* {11492}.
Type varieties	T. turgidum ssp. turgidum BGE048494 {11492}. Glu-B1cr [{11492}]. 8*.1+20y {11492}.
Type varieties	T. turgidum ssp. durum BGE045649 {11492}, BGE047535 {11492}. Glu-B1cs [{11492}]. 20x {11492}.
Type varieties	T. turgidum ssp. durum BGE045673 {11492}. Glu-B1ct [{11540}]. 6+(8) {11540}.
Type varieties	T. turgidum ssp. durum Langdon {11540}.

GLU-D1

Synonym

Glt-D2 {420}, Glu 1D {1415}, Glt-D1 {420}

1DL {150},{1125},{107},{780},{1100}.

Glu-D1a

Note	2+12 {1116}.
Varieties	CS. Primers were designated that enabled Dx2 to be distinguished from Dx5 and Dy10 from Dy12 {10641}.

Glu-D1b

Note	3+12 {1116}.
Varieties	Hobbit.

Glu-D1c

Note	4+12 {1116}.
Varieties	Champlein.

Glu-D1d

Note	5+10 {1116}.
Varieties	Hope.

Glu-D1e

Note	2+10 {1116}.
Varieties	Flinor (rare).

Glu-D1f

Note	2.2+12 {1116}.
Varieties	Danchi (rare). Glu-D1f is present at high frequencies in wheats of southern Japan. Its presence may be associated with white salted noodle (Udon) quality {10573}.

Glu-D1g

Note	5+9 {478}.
Varieties	BT-2288. Subunit 9 of Glu-D1g (5+9) was so numbered in {478} because its mobility is the same as one of the subunits encoded by Glu-B1c

Glu-D1h

Note	5+12 {1145}.
Varieties	Fiorello, Italy. Cultivar Fiorello is given as a standard for Glu-D1h encoding subunits 5+12 and for Glu-D1w encoding subunits 5[*] +10. An attempt to resolve

Glu-D1i

Note	Null {107}.
Varieties	Nap Hal, Nepal.

Glu-D1j

Note	2+12[*] {1146}.
Varieties	Tudest.

Glu-D1k

Note	2 {421}.
Sources / synonyms	CS/Timstein 1D. Glu-D1k {421} appears to have arisen as the result of a deficiency of subunit 12 from Glu-D1a (2+12); subunits 2 and 12 are

Glu-D1l

Note	12 {759}.
Varieties	AUS 10037, Afghanistan.

Glu-D1m

Note	10 {759}.
Varieties	AUS 13673, Afghanistan.

Glu-D1n

Note	2.1+10 {759}.
Varieties	AUS 14653, Afghanistan.

Glu-D1o

Note	2.1+13 {755}.
Varieties	AUS 14519, T. macha . One of the Glu-D1o subunits was numbered 13 in {755}; 13 was previously used to number a subunit encoded by

Glu-D1p

Note	36 {1233}.
i	Iranian landrace accession 3048/5[*] Sicco.

Glu-D1q

Note	2+11 {124}.
Varieties	Flinor.

Glu-D1r

Note	2.3+12 {1229}.
Varieties	PI 348465.

Glu-D1s

Note	38 {1032}.
Varieties	Leningradka.

Glu-D1t

Note	43+44 {668}.
dv	Ae. tauschii accession TA2450/2[*] .

Glu-D1u

Note	2+10' {836}.
Varieties	Coker 68-15.

Glu-D1v

Note	2.1+10.1 {755}.
dv	Ae. tauschii .

Glu-D1w

Note	5[*] +10 {03124}.
Varieties	Fiorello {03124}. Note that the cultivar Fiorello is given as a standard for Glu-D1h encoding subunits 5+12 and for GluD1w encoding subunits 5[*] +10. An

Glu-D1x

Note	2+T2 {755}.2[t] +12.2[t ] {03124}.
dv	Ae. tauschii .

Glu-D1y

Note	3[t] +12.2[t ] {03124}. 3+T2 {755}.
dv	Ae. tauschii .

Glu-D1z

Note	3+10 {755}.
dv	Ae. tauschii .

Glu-D1aa

Note	3+10.3 {755}.
dv	Ae. tauschii .

Glu-D1ab

Note	4.1+10 {755}.
dv	Ae. tauschii .

Glu-D1ac

Note	4+10 {755}.
dv	Ae. tauschii .

Glu-D1ad

Note	5.1+10.2 {755}.
dv	Ae. tauschii .

Glu-D1ae

Note	2.1[t] +12.2[t] {03124}. 2.1+T2 {1578}.
dv	Ae. tauschii .

Glu-D1af

Currently undesignated for reasons given in the preamble to this section.

Glu-D1ag

Note	1.5+T2 {1578}. 1.5[t] +12.2[t ] {03124}.
dv	Ae. tauschii .

Glu-D1ah

Note	1.5+10 {1578}.
dv	Ae. tauschii .

Glu-D1ai

Note	2.1+10.5 {1578}.
dv	Ae. tauschii .

Glu-D1aj

Note	1.5+12 {1578}.
dv	Ae. tauschii .

Glu-D1ak

Note	3+10.5 {1578}.
dv	Ae. tauschii .

Glu-D1al

Note	2.2[* ] {02107}.
Varieties	MG315.

Glu-D1am

Synonym

Glu-D1-I {03122}

2+12' {03122}.

Glu-D1an

Synonym

Glu-D1-II {03122}

2+12[* ] {03122}.

Glu-D1ao

Synonym

Glu-D1-III {03122}

2.4+12 {03122}.

Glu-D1ap

Synonym

Glu-D1-IV {03122}

2.5+12 {03122}.

Glu-D1aq

Note	1.5[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1ar

Note	2[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1as

Note	1.5[t] +10.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1at

Note	3[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1au

Note	2.1[t] +10.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1av

Note	2[t] +12.3[t] {03124}.
dv	Ae. tauschii .

Glu-D1aw

Note	1[t] +10[t] {03124}.
dv	Ae. tauschii .

Glu-D1ax

Note	1[t] +12[t] {03124}.
dv	Ae. tauschii .

Glu-D1ay

Note	1[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1az

Note	4[t] +12.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1ba

Note	1[t] +12.3[t] {03124}.
dv	Ae. tauschii .

Glu-D1bb

Note	1.5[t] +11[t] {03124}.
dv	Ae. tauschii .

Glu-D1bc

Note	1.5[t] +10.3[t ] {03124}.
dv	Ae. tauschii .

Glu-D1bd

Note	1[t] +11[t] {03124}.
dv	Ae. tauschii .

Glu-D1be

Note	2.1[t] +12.4[t] {3124}.
dv	Ae. tauschii .

Glu-D1bf

Note	2[t] +12.1[t] {03124}.
dv	Ae. tauschii {03124}.

Glu-D1bg

Note	3[t] +10.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1bh

Note	4[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1bi

Note	4[t] +10.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1bj

Note	5[t] +11[t] {03124}.
dv	Ae. tauschii .

Glu-D1bk

Note	5[t] +10.1[t] {03124}.
dv	Ae. tauschii .

Glu-D1bl

Note	5[t] +12.2[t] {03124}.
dv	Ae. tauschii .

Glu-D1bm

Note	5[*t] +null {03124}.
dv	Ae. tauschii .

Glu-D1bn

Note	5[*t] +12 {3124}.
dv	Ae. tauschii .

Glu-D1bo

Note	5'+12 {10091}.
Varieties	W958 {10091}. This putative new allele encodes two subunits that have very similar electrophoretic mobilities compared to subunits 5+12 encoded by Glu-D1h , but analysis

Glu-D1bp

Note	2.1'+12 {10327}.
Varieties	KU-1034 {10327}.

Glu-D1bq

Synonym

Glu-D1bp(t) {10304}

2.6+12 {10304}.

Glu-D1br

Note	5*t+10.1[t] {10426}.
Type varieties	Ae. tauschii TD81 {10426}. Subunit 10.1[t] possesses a mobility slightly lower than subunit 10 in SDS-PAGE and its deduced amino

acid sequence is similar to subunit 12 (8 amino acid differences) {10426}; the authors used the complete coding sequence to make phylogenetic comparisons with 19 other subunits including both x-type and y- type subunits and concluded that a Glu-1 gene duplication event occurred about 16.83 million years ago.

Glu-D1bs

Note	1.6[t] +12.3[t] {10642}.
dv	Ae. tauschii TD16 {10642}.

Glu-D1bt

Note	2.1[t] +12[t] {10568}.
Varieties	Syn 396{10568}.

Glu-D1bu

Note	2'+12 {10810}.
Varieties	CWI-64806, T. aestivum var. aestivum {10810}.

Glu-D1bv

Note	2''+10 {10810}.
Varieties	CWI-65297, T. aestivum var. erythroleucon {10810}.

Glu-D1bw

Note	2''+12 {10810}.
Varieties	CWI-60509, T. aestivum var. graecum {10810}.

GLU-Ag[i] 1

Note	1Ag[i] {374}.
ad	Vilmorin 27/ Th. intermedium .

GLU-E1

Chromosome	1ES
ad	CS/ E. elongata .

HMW glutenin y-type subunit Ee1.5 encoded by this locus was sequenced {10439} and compared with other y-type subunits, particularly subunit 1Dy10. It has major deletions in its middle region and is one of the smallest known HMW glutenin subunits. It has an additional Cys residue in the middle of the repetitive domain but lacks one Cys residue commonly found towards the end of this domain. These changes may influence inter- or intra-molecular disulphide bond formation. Four {10660, 10661} and 11 {10662} alleles were observed in Agropyron elongatum (E[e] genome, 2n = 10X = 70) and named Aex1 to Aex5 (producing x-type subunits) and Aey1 to Aey10 (producing y-type subunits). Aex4, Aey7 and Aey9 were very similar to three alleles in the diploid progenitor Lophopyrum elongatum {10439, 10663}. The C-terminal regions of three of the y-type subunits (products of Aey8, Aey9 and Aey10 ) were more similar to x-type subunits than to other y-type subunits {10662}. The subunit from Aex4 contained an additional cysteine residue, which may be associated with good processing quality in wheat introgression lines {10662}. Allele Aey-4 was a chimeric gene formed by recombination of two other genes {10662}. Chinese T. aestivum cultivar Xiaoyanmai carries a subunit with electrophoretic mobility in 10% SDSPAGE well beyond that of subunits so far observed in T. aestivum . It may derive from Agropyron elongatum , which was used in the breeding program that led to the variety {1538}. It has not been given a subunit number or allelic designation, because its genetic control has not been elucidated.

Glu-E1a

ad	CS/ L. elongatum W0622 {781}.

Glu-E1b

ad	Langdon/ L. elongatum DGE-1 {10644}.
al	L. elongatum PI 531719 {10644}.

GLU-H1

Synonym	Hor 3 {1337}
Chromosome	1HL
Chromosome	1H
ad	CS/Betzes {781}.
al	Various barley cultivars {1337}.

GLU-H[ch] 1

Chromosome	1H
ad	CS/ H. chilense .

38 accessions (natural populations) of Hordeum chilense carrying the following 10 subunits were used as the maternal parents of 121 lines of primary tritordeum, and evaluations for associations with breadmaking quality initiated {03114}. Subunits 1[Hch] , 2[Hch] and 3[Hch] were previously referred to as H[ch] a, H[ch] b and H[ch] c {03112}.

Glu-H[ch] 1a

Note	1[Hch] {03114}.
al	H. chilense accession H1 {03114}.

Glu-H[ch] 1b

Note	2[Hch] {03114}.
al	H. chilense accession H11 {03114}.

Glu-H[ch] 1c

Note	3[Hch] {03114}.
al	H. chilense accession H7 {03114}.

Glu-H[ch] 1d

Note	4[Hch] {03114}.
al	H. chilense accesion H16{03114}.

Glu-H[ch] 1e

Note	5[Hch] {03114}.
al	H. chilense accession H47 {03114}.

Glu-H[ch] 1f

Note	6[Hch] {03114}.
al	H. chilense accession H220 {03114}.

Glu-H[ch] 1g

Note	7[Hch] {03114}.
al	H. chilense accession H293 {03114}.

Glu-H[ch] 1h

Note	8[Hch] {03114}.
al	H. chilense accession H297 {03114}.

Glu-H[ch] 1i

Note	9[Hch] {03114}.
al	H. chilense accession H252 {03114}.

Glu-H[ch] 1j

Note	10[Hch] {0 3114}.
al	H. chilense accession H210 {03114}.

GLU-H[t] 1

Note	1H[t] L {1037}.
ad	CS/ E. trachycaulum .

GLU-R1

Synonym	Sec 3 {1336}
Chromosome	1RL
Chromosome	1R
ad	CS/Imperial; Holdfast/ King II {1340}.
tr	CS Imperial 1DS.1RL {1356}.

Glu-R1a

Note	1[r] -4[r] {03116}.
Varieties	Indiana hexaploid triticale {03116}.

Glu-R1b

Note	2[r] -6.5[r] {03116}.
Varieties	Graal hexaploid triticale {03116}.

Glu-R1c

Note	6[r] -13[r] {03116}.
Varieties	Almao hexaploid triticale {03116}.

Glu-R1d

Note	2[r] -9[r] {03116}.
Varieties	Olympus hexaploid triticale {03116}.

Glu-R1e

Note	6.5[r] {03116}.
Varieties	Clercal hexaploid triticale {03116}.

Glu-R1f

Note	0.8[r] -6[r] {03115}.
Varieties	Carmara hexaploid triticale {03115}.

Glu-R1g

Note	5.8[r] {03115}.
Varieties	Arrayan hexaploid triticale {03115}. From study of chromosome substitutions in bread wheat {03117}, it was found that a chromosome 1R carrying HMW secalin subunit 6.5[r]
ad	CS/ S. montanum {1340, 1339}.

GLU-S[l] 1

Note	1S[l] L {1228}.
Marker associations	In Ae. longissima 2 /Ae. longissima 10, GLU-S[l] 1, G LU - S[l] 3 , one glucose phosphate isomerase locus, and three gliadin loci were

GLU-Ta1

al	Taenitherum crinitum PI 204577 {10449}.

Glu-A1-1a

Note	Null.
Varieties	CS.

Glu-A1-1b

Note	1.
Varieties	Hope.

Glu-A1-1c

Note	2[*] .
Varieties	Bezostaya 1. A PCR marker specific for the Glu-A1-1c (Ax2[*] ) allele was developed in {0147}.

Glu-A1-1d

Varieties

V74, Spain.

Glu-A1-1e

Varieties

132c, Poland.

Glu-A1-1f

Varieties

112-29, Sudan.

Glu-A1-1g

Varieties

Landrace 1600.

Glu-A1-1h

Type varieties

PI 94683, USSR, T. dicoccum .

Glu-A1-1i

Type varieties

CI 12213, India, T. dicoccum .

Glu-A1-1j

Note	1'.
Type varieties	PI 352359, Germany, T. dicoccum ; Lambro.

Glu-A1-1k

Note	26.
Varieties	BT-2288

Glu-A1-1l

Type varieties

Chinook, Canada.

Glu-A1-1m

Type varieties

Nugget Biotype 1, Canada.

Glu-A1-1n

Note	1".
Type varieties	Corado, Portugal.

Glu-A1-1o

Note	2[**] .
Type varieties	PI 61189, USSR, Aric 581/1.

Glu-A1-1p

Note	3[*] .
Varieties	David 1.

Glu-A1-1q

Note	2[***] .
Type varieties	Melianopus 1528.

Glu-A1-1r

Note	39.
i	T. thaoudar IPSR 1020006/6[*] Sicco.

Glu-A1-1s

Note	41.
i	T. thaoudar G3152/6[*] Sicco.

Glu-A1-1t

Note	21[* ] {602}.
Varieties	W29323, W 3879, W 31169. Glu-A1-1t is a provisional designation; definitive evidence that subunit 21[*] , which has a mobility similar to that of subunit

Glu-A1-1u

Note	2[*B] {02106}.
Varieties	Bankuti 1201.

Glu-A1-1v

Note	2.1* {10327}.
Varieties	Grado {10327}; KU-1026 {10327}; KU-1086 {10327}; KU1094 {10327}; KU-1139 {10327}.

Glu-A1-1w

Note	2' {10327}.
Varieties	TRI14165/91 {10327}.

Glu-A1-1x

2'' {10535}. **v:

Glu-A1-2a

Note	Null.
Varieties	CS.

Glu-A1-2b

Note	40.
i	T. thaoudar IPSR1020006/6[*] Sicco.

Glu-A1-2c

Note	42.
i	T. thaoudar G3152/6[*] Sicco.

Glu-B1-1a

Note	7.
Varieties	CS. A PCR marker (2373 bp) for the Glu-B1-1a (Bx7) allele was developed in {0145}.

Glu-B1-1b

Note	7,7[*] .
Varieties	Flinor, Bezostaya 1, Owens, Norstar.

Glu-B1-1c

Note	7'.
Varieties	Adonis.

Glu-B1-1d

Note	6.
Varieties	Hope.

Glu-B1-1e

Note	20.
Varieties	Federation.

Glu-B1-1f

Note	13.
Varieties	Lancota.

Glu-B1-1g

Note	14.
Varieties	Sappo.

Glu-B1-1h

Note	17.
Varieties	Gabo.

Glu-B1-1i

Note	21.21x {03116}.
Varieties	Dunav; Foison {03116}.

Glu-B1-1j

Note	23.
Varieties	Spica D.

Glu-B1-1k

Type varieties

PI 94640, Iran, T. dicoccum .

Glu-B1-1l

Type varieties

PI 355505, Germany, T. diccocum .

Glu-B1-1m

Type varieties

PI 352354, Ethiopia, T. dicoccum .

Glu-B1-1n

Type varieties

PI 94633, Morocco, T. dicoccum .

Glu-B1-1o

Varieties

Supreza, Canada.

Glu-B1-1p

Varieties

Mondor.

Glu-B1-1q

Type varieties

Canoco de Grao Escuro, Portugal.

Glu-B1-1r

Type varieties

Tremez Mollez, Portugal.

43 P ROTEINS

Glu-B1-1s

Type varieties

Quaduro, Italy.

Glu-B1-1t

Type varieties

Athena, Italy.

Glu-B1-1u

Note	26.
Varieties	Cologna 1.

Glu-B1-1v

Note	28.
Varieties	Forlani.

Glu-B1-1w

Note	Null.
Varieties	Olympic mutant.

Glu-B1-1x

Note	30.
Varieties	Marinar.

Glu-B1-1y

Note	32.
Varieties	BG-1943.

Glu-B1-1z

Note	34.
Varieties	Jeja Almendros.

Glu-B1-1aa

Note	37.
Varieties	Shedraya Polesja.

Glu-B1-1ab

Note	6[*] .
Varieties	Dawbill.

Glu-B1-1ac

Note	6.8 {03116}.
Varieties	Carnac hexaploid triticale {03116}.

Glu-B1-1ad

Note	13[*] {03122}.
Varieties	PI 348767 spelt {03122}.

Glu-B1-1ae

Note	14* {10327}.
Varieties	TRI11553/92 {10327}.

Glu-B1-1af

Note	6.1 {10327}.
Varieties	Hercule {10327}; KU-3418 {10327}; KU-3446 {10327}; Rouguin {10327}; Schwabenkorn {10327}; SP3 {10327}; Steiners Roter Tiroler {10327}; TRI4613/75 {10327}.

Glu-B1-1ag

Note	7[**] {10643}.
Varieties	XM1368-2 {10643}.

Glu-B1-1ah

Note	7[OE] {899}.
Varieties	Benkuti 1201 {10196, 10197}; Glenlea {899}; Klein Universal II {10196}; Tezanos Pintos Precoz {10196}; Tobari {10196}.

Glu-B1-1ai

Note	7.1 {10807}.
Varieties	H45 {10807}. Glu-B1-1aj 10807}. 7.2 {10807}.
Varieties	H45{10807}.

Glu-B1-1ak

Note	7.3 {10807}.
Varieties	VQ0437 {10807}.

Glu-B1-1al

Note	17' {10809}.
Type varieties	TGR-214 {10809}; TGR-2246 {10809}.

Glu-B1-1am

Note	13[**] {10809}.
Type varieties	TGR-003 {10809}.

Glu-B1-1an

Note	7b[*] {10808}.
Varieties	Eshimashinriki {10808}.

Glu-B1-1ao

Note	20* {11490}.
Type varieties	T. turgidum ssp. durum Mexican landrace accession 22 (CWI52215) {11490}.

Glu-B1-2a

Note	8.
Varieties	CS.

Glu-B1-2b

Note	9.
Varieties	Bezostaya 1.

Glu-B1-2c

Note	16.
Varieties	Lancota.

Glu-B1-2d

Note	19.
Varieties	NS 335.

Glu-B1-2e

Note	15.
Varieties	Sappo.

Glu-B1-2f

Note	18.
Varieties	Gabo.

Glu-B1-2g

Note	22.
Varieties	Serbian.

Glu-B1-2h

Note	24.
Varieties	Spica D

Glu-B1-2i

Type varieties

PI 355505, Germany, T. dicoccum .

Glu-B1-2j

Type varieties

PI 352354, Ethiopia, T. dicoccum .

Glu-B1-2k

Type varieties

PI 94633, Morocco, T. dicoccum .

Glu-B1-2l

Note	11.
Varieties	BT-2288.

Glu-B1-2m

Varieties

Supreza, Canada.

Glu-B1-2n

Varieties

Mondor.

Glu-B1-2o

Note	8[*] .
Varieties	Dawbull.

Glu-B1-2p

Type varieties

Canoco de Grao Escuro, Portugal.

Glu-B1-2q

Type varieties

Tremez Mollez, Portugal, T. durum .

Glu1-2r

Type varieties

Quaduro, Italy, T. durum .

Glu-B1-2s

Note	18[*] .
Varieties	David.

Glu-B1-2t

Note	27.
Varieties	Cologna 1.

Glu-B1-2u

Note	29.
Varieties	Forlani.

Glu-B1-2v

Note	Null.
Varieties	Olympic mutant.

Glu-B1-2w

Note	31.
Varieties	Marinar.

Glu-B1-2x

Note	33.
Varieties	BG-1943.

Glu-B1-2y

Note	35.
Varieties	Jeja Almendros.

Glu-B1-2z

Note	20y {03116}.
Varieties	Carnac hexaploid triticale {03116}.

Glu-B1-2aa

Note	18' {03122}.
Varieties	PI 348631 spelt {03122}.

Glu-B1-2ab

Note	21y {03116}.
Varieties	Foison {03116}.

Glu-B1-2ac

Note	22* {10327}.
Varieties	Grado {10327}; KU-1026 {10327}; KU-1086 {10327}; KU1094 {10327}; KU-1139 {10327}.

Glu-B1-2ad

Note	22.1 {10327}.
Varieties	Hercule {10327}; KU-1135 {10327}; Rouguin {10327}; Schwabenkorn {10327}; SP3 {10327}; Steiners Roter Tiroler {10327}.

Glu-B1-2ae

Note	15* {10327}.
Varieties	TRI11553/92 {10327}.

Glu-B1-2af

Note	19* {10327}.
Varieties	KU-3410 {10327}; Rechenbergs Fruher Dinkel {10327}; Renval {10327}; SP1{10327}; TRI9885/74{10327}; Zeiners Weiser Schlegel {10327}.

Glu-B1-2ag

Synonym	8[**] {10643}
Varieties	XM1404-2 {10643}. Glu-B1-2ah . Currently undesignated.

Glu-B1-2ai

Note	8' {10809}.
Type varieties	TGR-244 {10809}.

Glu-B1-2aj

Note	8a[*] {10808}.
Varieties	Jing 411{10808}; Pioneer {10808}; Tasman {10808}.

Glu-B1-2ak

Note	8b[*] {10808}.
Varieties	ACA 601 {10808}; Nidera Baguette 10 {10808

Glu-B1-2al

Note	33* {11490}.
Type varieties	T. turgidum ssp. durum Iranian landrace accession 51 (CWI57280) {11490}.

Glu-B1-2am

Note	22* {11490}.
Type varieties	T. turgidum ssp. durum Iranian landrace accession 51 (CWI57280) {11490}. Glu-B1-2an [{11492}]. 8*.1 {11492}.
Type varieties	T. turgidum ssp. durum BGE045649 {11492}, BGE047535 {11492}. Glu-B1-2ao [{11540}]. (8) {11540}.
Type varieties	T. turgidum ssp. durum Langdon {11540}.

Eight alleles at GLU-B1-1 and 10 alleles at GLU-B1-2 in T. turgidum var. dicoccoides populations were described in {798}. In a further study using different germplasm of this species {205}, 19 alleles at GLUB1 were observed, including 15 not previously observed; the 19 alleles included 11 alleles at GLU-B1-1 and 14 alleles (including the null allele) at GLU-B1-2 , although, as the authors pointed out, it was not conclusively clear how many of these alleles were distinct from each other, or from others previously observed.

GLU-D1-1 Glu-D1-1a

Note	2.
Varieties	CS.

Glu-D1-1b

Note	3.
Varieties	Hobbit.

Glu-D1-1c

Note	4.
Varieties	Champlein.

Glu-D1-1d

Note	5.
Varieties	Hope. PCR markers specific for the Glu-D1-1d (Dx5) allele were developed in {0145} and {0147}.

Glu-D1-1e

Note	2.2.
Varieties	Danchi.

Glu-D1-1f

Note	Null.
Varieties	Nap Hal, Nepal.

Glu-D1-1g

Note	2.1.
Varieties	AUS 14653, Afghanistan.

Glu-D1-1h

Note	2.3.
Varieties	PI 348465.

Glu-D1-1i

Note	38.
Varieties	Leningradka.

Glu-D1-1j

Note	43 {668}.
i	Ae. tauschii accession TA2450/2[*] .

Glu-D1-1k

Note	4.1 {755}.
dv	Ae. tauschii .

Glu-D1-1l

Note	1.5 {1578}. D[t] x1.5 {10306}.
dv	Ae. tauschii accession SQ-214 {10306}. A restriction enzyme-based method named the 'restricted deletion method' was used to characterize the ORF of this subunit {10306} (as

Glu-D1-1m

Note	2.2[*] {02107}.
Varieties	MG315.

Glu-D1-1n

Note	2.4 {03122}.
Varieties	PI 348473 spelt {03122}.

Glu-D1-1o

Note	2.5 {03122}.
Varieties	PI 3484572 spelt {03122}.

Glu-D1-1p

Note	1[t] {03124}.
dv	Ae. tauschii {03124}.

Glu-D1-1q

Note	5[*t] {03124}.
dv	Ae. tauschii {03124}.

Glu-D1-1r

Note	5.1 {755}.
dv	Ae. tauschii . This allele was designated Glu-D1-1j in the 1998 Catalogue edition.

Glu-D1-1s

Note	5' {10091}.
Varieties	W958 {10091}. This putative allele encodes a subunit, provisionally denominated 5' {10091}, that has a very similar electrophoretic mobility compared to subunit 5 encoded by

Glu-D1-1t

Note	2.6 {10304}.
Varieties	Baidongmai {10305}; Jinbaojin {10305}; Hongdongmai {10305}; Hongkedongmai {10305}.

Glu-D1-1u

Note	2.1' {10327}.
Varieties	KU-1034{10327}.

Glu-D1-1v

Synonym	1.6[t ] {10642}
dv	Ae. tauschii TD16 {10642}.

GLU-D1-2 Glu-D1-2a

Note	12.
Varieties	CS. A PCR marker (612 bp) for the Glu-D1-2a (Dy12) allele was developed in {0145}.

Glu-D1-2b

Note	10.
Varieties	Hope. PCR markers (576 bp and 2176 bp) for the Glu-D1-2b (Dy10) allele were developed in {0145} and {0147}, respectively.

Glu-D1-2c

Note	9.
Varieties	BT-2288.

Glu-D1-2d

Note	Null.
Varieties	Nap Hal, Nepal.

Glu-D1-2e

Note	12[*] .
Varieties	Tudest.

Glu-D1-2f

Note	13.
Varieties	AUS 14519, T. macha .

Glu-D1-2g

Note	36.
i	Iranian landrace 3048/5[*] Sicco.

Glu-D1-2h

Note	11.
Varieties	Flinor.

Glu-D1-2i

Note	44 {668}.
i	Ae. tauschii TA2450/2[*] .

Glu-D1-2j

Note	10' {836}.
Varieties	Coker 68-15.

Glu-D1-2k

Note	T1 {755}.
dv	Ae. tauschii .

Glu-D1-2l

Note	T2 {755}.
dv	Ae. tauschii .

Glu-D1-2m

Note	10.1 {755}.
dv	Ae. tauschii .

Glu-D1-2n

Note	10.2 {755}.
dv	Ae. tauschii .

Glu-D1-2o

Note	10.3 {755}.
dv	Ae. tauschii .

Glu-D1-2p

Note	10.5 {1578}.
dv	Ae. tauschii .

Glu-D1-2q

Note	12' {03122}.
Varieties	PI-348495 spelt wheat accession {03122}.

Glu-D1-2r

Note	12.1[t ] {03124}.
dv	Ae. tauschii .

Glu-D1-2s

Note	12.3[t] {03124}.
dv	Ae. tauschii .

Glu-D1-2t

Note	12.4[t] {03124}.
dv	Ae. tauschii .

Glu-D1-2u

Note	D[t] y10 {10306}.
Varieties	Ae. tauschii accession SQ-214 {10306}. A restriction enzyme-based method named the 'restricted deletion method' was used to characterize the

ORF of this subunit {10306} (as in the case of subunit 1.5 (or D[t] x1.5 {10306}) encoded by Glu-D1-1l {10306}. This subunit was first recognized as being different from subunit 1- encoded by Glu-D1-2b in hexaploid wheat in {10307}.Six combinations involving 5 HMW subunits [1A (u-z)] are listed in {420}, from a study of 109 genotypes including representatives of botanical varieties. Alleles in T. turgidum var. dicoccoides populations, 12 at GLU-A1-1 and 3 at GLU-A1-2 , were described in {798}. In a further study using different germplasm of this species {205}, 14 alleles at GLU-A1 were observed, including 12 not previously found; the 15 alleles included up to 15 alleles at GLU-A1-1 (with up to 10 not previously observed), and 5 alleles at GLU-A1-2 (with 4 not previously observed) (numbers take the null allele into account). The uncertainty in numbers is due to the very similar electrophoretic mobilities of some of the subunits compared with others observed either in this study or previously. In a study including emmers ( T. dicoccum ) {00115}, new subunits named 1[+] and 2[-] were found in accessions MG4378/1 and MG5380/1, respectively, and provisionally assigned to GLU-A1 . Until confirmed, they are not included in the GLU-A1 list.

Glu-R1-1a

Note	1[r] {03116}.
Varieties	Indiana hexaploid triticale {03116}.

Glu-R1-1b

Note	2[r] {03116}.
Varieties	Graal hexaploid triticale {03116}.

Glu-R1-1c

Note	6[r] {03116}.
Varieties	Alamo hexaploid triticale {03116}.

Glu-R1-1d

Note	0.8[r] {03115}.
Varieties	Carmara hexaploid triticale {03115}.

Glu-R1-1e

Note	5.8[r] {03115}.
Varieties	Arrayan hexaploid triticale {03115}.

Glu-R1-2

Note	1R,
Chromosome	1RL

Glu-R1-2a

Note	4[r] {03116}.
Varieties	Indiana hexaploid triticale {03116}.

Glu-R1-2b

Note	6.5[r] {03116}.
Varieties	Graal hexaploid triticale {03116}.

Glu-R1-2c

Note	13[r] {03116}.
Varieties	Alamo hexaploid triticale {03116}.

Glu-R1-2d

Note	9[r] {03116}.
Varieties	Olympus hexaploid triticale {03116}. There was difficulty in assigning subunit 6[r] in the GLU-R1-1 and GLU-R1-2 lists, since it appeared as an x-type subunit in

GLU-V1-1

Alleles and subunits at GLU-V1-1 and GLU-V1-2 : The following is analogous to the GLU-1-1 and GLU1-2 lists given earlier to identify x-type and y-type subunits in wheat. It was assumed that where an allele at GLU-V1 produces only a single subunit, it is an x-type subunit and so encoded by GLU-V1-1 rather than by GLU-V1-2 ; the electrophoretic mobilities of the subunits are all greater, though some only marginally so, than subunit 7 encoded by Glu-B1-1a (an x-type subunit), and extend beyond the mobility of subunit 12 encoded by Glu-D1-2a (a y-type subunit) {1651}; therefore, it is quite possible that any one of the subunits designated as encoded by GLU-V1-1 is, in fact, encoded by GLU-V1-2 . The designation given here is intended to be the most practically useful until the identities of the genes encoding the alleles are directly established.

Glu-V1-1a

Note	71 {1651}.
al	D. villosum .

Glu-V1-1b

Note	72 {1651}.
al	D. villosum .

Glu-V1-1c

Note	73 {1651}.
al	D. villosum .

Glu-V1-1d

Note	74 {1651}.
al	D. villosum .

Glu-V1-1e

Note	75 {1651}.
al	D. villosum .

Glu-V1-1f

Note	76 {1651}.
al	D. villosum .

Glu-V1-1g

Note	77 {1651}.
al	D. villosum .

Glu-V1-1h

Note	78 {1651}.
al	D. villosum .

Glu-V1-1i

Note	79 {1651}.
al	D. villosum .

Glu-V1-1j

Note	80 {1651}.
al	D. villosum .

Glu-V1-1k

Note	Null {1651}.
al	D. villosum .

Glu-V1-1l

Note	81 {1651}.
al	D. villosum .

Glu-V1-1m

Note	83 {1651}.
al	D. villosum .

Glu-V1-1n

Note	85 {1651}.
al	D. villosum .

GLU-V1-2 Glu-V1-2a

Note	Null {1651}.
al	D. villosum

Glu-V1-2b

Note	82 {1651}.
al	D. villosum .

Glu-V1-2c

Note	84 {1651}.
al	D. villosum .

Glu-V1-2d

Note	86 {1651}.
al	D. villosum .

TRI-A1

Chromosome	1AS
Varieties	CS.

Tri-A1a

Synonym	cs {1358}
Varieties	CS.

Tri-A1b

Synonym	h {1358}
Varieties	Hope.

TRI-D1 {707}, {1358}, {1357}. 1DS {1357}.

Tri-D1a

Synonym	cs {1358}
Varieties	CS.

Tri-D1b

Synonym	i {1358}
Varieties	India 115.

ISA1 ISA-A1

Chromosome	2AL
Varieties	CS.

Isa-A1a

Varieties

CS.

Isa-A1b

Note	Null allele.
Varieties	Cajeme 71.

ISA-B1

Chromosome	2BL
Varieties	CS.

Isa-B1a

Varieties

CS.

Isa-B1b

Varieties

Bihar.

ISA-D1

Chromosome	2DL
Varieties	CS.

Orthologous genes were identified in Ae. speltoides and T. timopheevii {908}. All durum wheats investigated had the genotype Isa-A1b, Isa-B1b .

GSP-A1

Synonym

GSP {614}

5A {614}, {383}.

GSP-B1

Synonym	GSP {614}
Chromosome	5B
Varieties	CS {614}; Glenlea {0385}. In {1185} sequence of clone TSF33 from cv. Soft Falcon (GenBank X80379) was identical to this allele, as are ESTs

GSP-D1

Synonym	GSP {614}
Chromosome	5DS

Gsp-D1a

Varieties	CS {614}; Glenlea {0385}.
dv	Ae. tauschii CPI1110799 (GenBank AF177219) {0383}.
Marker associations	Co-segregation of Gsp-D1 and Ha {614}.

Gsp-D1b

dv	Ae. tauschii TA1583 (GenBank AY252079) Pina-D1a, Pinb-D1a {3105}; TA2475 (GenBank AY252087) Pina-D1a, Pina-D1i {03105}.

Gsp-D1c

dv	Ae. tauschii TA2369 (GenBank AY252081) Pina-D1c, Pinb-D1h {03105}; CPI110799 (GenBank AF177219) {0383}.

Gsp-D1d

dv	Ae. tauschii TA2536 (GenBank 252093) Pina-D1c, Pinb-D2i {03105}; TA2374 (GenBank AY252046) Pina-D1d, Pinb-D1i {03105}; TA2458 (GenBank AY252084) Pina-D1e, PinbD1i {03105}; TA2436 (GenBank

Gsp-D1e

dv	Ae. tauschii TA2527 (GenBank AY252066) Pina-D1c, Pinb-D1h {03105}; TA2512 (GenBank AY252092) Pina-D1d, Pinb-D1i {03105}; TA2495 (GenBank AY252091) Pina-D1e, PinbD1i {03105}.

Gsp-D1f

dv	Ae. tauschii TA1649 (GenBank AY252063) Pina-D1d, Pinb-D1h {03105}; TA2455 (GenBank AY252073) Pina-D1d, Pinb-D1i {03105}.

Gsp-D1g

dv	Ae. tauschii TA1599 (GenBank AY252062) Pina-D1a, Pinb-D1j {03105}.

Gsp-D1h

dv	Ae. tauschii TA1691 (GenBank AY252064) Pina-D1a, Pinb-D1j {03105}.

Gsp-D1i

Varieties

Yecora Rojo (GenBank AY255771) Pina-D1b, Pinb-D1a {03105}.

Gsp-D1j

Sources / synonyms

CS*/Red Egyptian 5D, Pina-D1, Pinb-D1 and Gsp-D1 {10077}. In {1185} the sequence of clone TSF69 from cv. Soft Falcon (GenBank S72696) is identical, as are

HSTH1-1 HSTH1-A1

Chromosome	5AL
Varieties	CS {0215}.

HSTH1-B1

Chromosome	5BL
Varieties	CS {0215}.

HSTH1-D1

Chromosome	5DL
Varieties	CS {0215}.

HSTH1-2 HSTH1-A2

Chromosome	5AL
Varieties	CS {0215}.

HstH1-A2a

Varieties

CS {0215}.

HstH1-A2b

Note	Null allele {0215}.
Varieties	Mara {0215}; 10 others{0215}.

HSTH1-B2

Chromosome	5BL
Varieties	CS {0215}.

HstH1-B2a

Varieties

CS {0215}; 19 others {0215}.

HstH1-B2b

Varieties

Excelsior {0215}.

HSTH1-D2

Chromosome	5DL
Varieties	CS {0215}.

HstH1-D1a

Varieties

CS {0215}; 18 others {0215}.

HstH1-D1b

Varieties

Grekum 114 {0215}; Kirgizsky Karlik {0215}.

The relationship of this gene series with a Hst-A1, Hst-B1, Hst-D1 series in group 5 chromosomes {0216} based on DNA hybridization studies was not established.

IBF-1 IBF-A1

Chromosome	5AL
Varieties	CS.

Ibf-A1a

Varieties

CS.

Ibf-A1b

Varieties

Cappelle-Desprez.

Ibf-A1c

Varieties

Hope.

Ibf-A1d

Varieties

Chris.

Ibf-A1e

Varieties

Sears' Synthetic.

IBF-B1

Chromosome	5BL
Varieties	CS.

Ibf-B1a

Varieties

CS.

Ibf-B1b

Varieties

Cappelle-Desprez.

Ibf-B1c

Varieties

Ciano 67.

Ibf-B1d

Varieties

Sears' Synthetic.

IBF-D1

Chromosome	5DL
Varieties	CS.

Ibf-D1a

Varieties

CS.

Ibf-D1b

Varieties

Cappelle-Desprez.

Ibf-D1c

Varieties

Purple Pericarp.

Ibf-D1d

Varieties

Sears' Synthetic.

IBF-Ag [i]

Note	5Ag[i] {818}.
ad	Vilmorin/ Th. intermedium .

IBF-E1

Chromosome	5EL
ad	CS/ E. elongata .

IBF-H1

Chromosome	4H
ad	CS/Betzes.

IBF-R1

Chromosome	5RL
ad	CS/Imperial, CS/KingII.

IBF-S[l] 1

Chromosome	5S
ad	CS/ Ae. sharonensis .

IBF-U1

Chromosome	5U
ad	CS/ Ae. umbellulata .

PUR-A1

Chromosome	1AL
Varieties	CS {351}. A PCR marker specific for PUR-A1 was developed in {9976}.

PUR-B1

Chromosome	1BL
Varieties	CS {351}. A PCR marker specific for PUR-B1 was developed in {9976}.

PUR-D1

Chromosome	1DL
Varieties	CS {351}. PCR marker specific for PUR-D1 was developed in {9976}. A locus in chromosome 5DS affects the level of lipopurothionin {351}.

PUR-R1

Note	1RL {1261} =
Chromosome	1RS
Chromosome	1BL
ad	CS/Imperial.
su	Several 1R(1B) lines.
tr	Aurora, Kavkaz. A PCR marker specific for PUR-R1 was developed in {9976}.

LEC-1 LEC-A1

Chromosome	1AL
Varieties	CS.

LEC-B1

Chromosome	1B
Sources / synonyms	CS[*] /Hope 1B.

LEC-D1

Chromosome	1DL
Varieties	CS.

LEC-U1

Chromosome	1U
ad	CS/ Ae. umbellulata .

PINa-A1

dv	T. urartu unspecified accession {03103}; TA763 (GenBank AJ302094) {03104, 03108}; TA808 (GenBank AJ302095) {03104, 03108}.

PINa-A[m] 1

Note	5A[m] S {0083}.
dv	T. monococcum DV92 (cultivated), G3116 (spp. aegilopoides ) (GenBank AJ242715) {0083}; unspecified acession (GenBank AJ249933) {03103}; PI277138 (GenBank AJ302093) {03104}; PI418582 (GenBank AJ302092)

TA183, TA291, TA546, TA581 (GenBank AY622786) {03108}. In T. monococcum PINa-A[m] 1 is completely linked to GSP-A[m] 1 {0083}.

PINa-D1

Chromosome	5DS
Varieties	CS (GenBank DQ363911) {03108}; Capitole (GenBank X69914) {03110}. This locus has a large deletion encompassing genes PINa-D1, PINb-D1 and GSP-D1 . This allelic combination

Pina-D1a

Varieties	Bellevue {0249}; Capitole (GenBank X69914) {03110}; Courtot {0249}; Fortuna {0249}; Galaxie {0249}; Heron {1035}; Renan (GenBank CR626934) {10440};
Varieties (alt.)	Aurelio Pinb-D1a {0249}; Bezostaja Pinb-D1b {0249}; Bilancia Pinb-D1a {0249}; Bolero Pinb-D1a {0249}; Brasilia Pinb-D1b {0249}; Centauro Pinb-D1a {0249}; Cerere
dv	Ae. tauschii upspecified accession (GenBank AJ249935) {03103}; TA2475 (GenBank AY252037) PinbD1i, Gsp-D1b {03105}; TA1599 (GenBank AY252011) Pinb-D1j, Gsp-D1g {03105}; TA1691 (GanBank AY252013)

Pina-D1a is present in all soft hexaploid wheats and possibly all hard hexaploid wheats that carry a hardness mutation in puroindoline b {452}, {1035}, {0082}, {0204}, {0295}.

Pina-D1b

Note	Null allele.
i	Falcon/7[] Heron, Heron/7[] Falcon {03109}; Gamenya Seln.{0203, 0298}; Heron/7[] Falcon sel. {0203, 0298}; PI 644080 (Alpowa/ID377s//7Alpowa) {10429}; Nearisogenic pairs were developed in
Varieties	Butte 86 {1035}; Eridano {0249}; Falcon {1035}; Glenlea (GenBank AB262660). This BAC clone also contains Pinb-D1a {10431}; Kalyansona{0249}; Super X
Varieties (alt.)	Amidon Pinb-D1a {0249}; Ciano Pinb-D1a {0249}; Dorico Pinb-D1a {0249}; Golia Pinb-D1a {0249}; Guadalupe PinbD1a {0249}; Barra Pinb-D1a {0249}; Inia

Present only in some hard hexaploid wheats. Pina-D1b is associated with harder texture than Pinb-D1b {0177, 0206}. This allele is now defined as a 15,380 bp deletion versus other possible puroindoline a nulls {10428, 10391}.

Pina-D1c

dv	Ae. tauschii TA2369 (GenBank AY252031) Pinb-D1h, Gsp-D1c ; TA2527 (GenBank AY252015) Pinb-D1h, Gsp-D1e {03108}; Ae. tauschii TA10 (GenBank AY649746) {03108}.

Pina-D1d

dv	Ae. tauschii PI452131 (GenBank AJ302098) Pinb-D1i {03104}; PI554318 (GenBank AJ302099) Pinb-D1k {03104}; TA1649 (GenBank AY252012) Pinb-D1h, Gsp-D1f {03105}; TA2374 (GenBank AY251996) Pinb-D1i, Gsp-D1d

Pina-D1e

dv	Ae. tauschii TA2458 (GenBank AY252034) Pinb-D1i, Gsp-D1d {03105}; TA2495 (GenBank AY252041) Pinb-D1i, Gsp-D1e {03105}.

Pina-D1f

dv	Ae. tauschii TA2436 (GenBank AY251998) Pinb-D1i, Gsp-D1d {03105}.

Pina-D1g

dv	Ae. tauschii TA1583 (GenBank AY252029) Pinb-D1a, Gsp-D1b {03105}.

Pina-D1h

Varieties

X. aegilotriticum CIGM86.946-1B-0B-0PR-0B (GenBank AY573898) Pinb-D1o {10118}.

Pina-D1i

Varieties

X. aegilotriticum CIGM87.2784-1B-0PR-0B (GenBank AY573899) Pinb-D1k {10118}.

Pina-D1j

Varieties

X. aegilotriticum CIGM88.1363-0B (GenBank AY573900) Pinb-D1o {10118}.

Pina-D1k

Synonym	homonym: Pina-D1b/Pinb-D1h(t)
Sources / synonyms	CS/Red Egyptian 5D substitution line, Pinb-D1q, Gsp-D1i* {10077}.
Varieties	Bindokku {10305}; Cheyenne-A {10305}; Chosen 68 {10305}; Gaiyuerui {10316}; KT020-584 {10432}; Saiiku 18 {10305}; Saiiku 44 {10305}; Safangmai

Pina-D1l

Synonym	Pina-D1c {10168}
Varieties	Baikezaomai Chinese landraces {10208}; Chengduguangtou {10208}; Guangtouxiaomai {10208}; Sanyuehuang {10208}; Xiaoyuhua {10208}.
Varieties (alt.)	Fortuna (USA) Pinb-D1a {10168}; Glenman Pinb-D1a {10168}. Pina-D1l has a C deletion leading to an open reading frame shift and premature stop codon; PINA

Pina-D1m

Varieties

Hongheshang (GenBank EF620907) {10208}. C-to-T substitution: Proline-35 to serine; hard kernel texture {10208}.

Pina-D1n

Varieties

Baimangchun {10208}; Hongheshang (GenBank EF620907) {10208}; Xianmai (GenBank EF620908) {10208}; Yazuixiaomai Chinese landraces {10208}; Yazuizi {10208}; Zhuantoubaike {10208}. G-to-A substitution: Tryptophan-43

Pina-D1o

dv	Ae. tauschii RM0182 (GenBank AY608595) {10311}.

Pina-D1p

Varieties

T. aestivum Jing 771 (GenBank AY599893) {10316}.

Pina-D1q

Varieties

U29 (GenBank AB181238) {10316}; Muu-27 (homonym 'a2', Pina-D1p ) {10316}.

PINb-A1

dv	T. urartu TA763 (GenBank AJ302103) {3104}; TA808 (GenBank AJ302104) {03104, 03108}.

Pinb-D1a

Varieties	Hill 81 {452}.
Varieties (alt.)	Adder Pina-D1a {0317}; Amidon Pina-D1b {0249}; Aurelio Pina-D1a {0249}; Barra Pina-D1b {0249}; Bilancia Pina-D1a {0249}; Bolero Pina-D1a {0249}; Centauro Pina-D1a
dv	Ae. tauschii unspecified accession (GenBank AJ249936) {03103}; TA1583 (GenBank AY251981) PinaD1a, Gsp-D1b {03105}.

Pinb-D1a is present in all soft hexaploid wheats and possibly all hard hexaploid wheats carrying the PinbD1b, -D1c, -D1d, -D1e, or -D1f mutations {452}, {1035}, {0082}, {0204}, {0295}.

Pinb-D1b

Chromosome	5DS
i	Paha[] 2/Early Blackhull/5[] Paha {0203,0298}; Early Blackhull der./5[] Nugaines seln. {0203, 0298}; hard sib sel. from Weston {03107}; PI 644081 (Alpowa/ND2603//7Alpowa) {10429}.
Sources / synonyms	CS[*] 7/Cheyenne 5D {452}.
Varieties	Thatcher {0204}; Wanser {452}; hard component of Turkey {0204}; Cheyenne (GenBank DQ363914) {10315}; Renan

(GenBank CR626934) {10440}.

Pinb-D1c

i	PI 644082 (Alpowa/Red Bobs//7*Alpowa) {10429}.
Varieties	Avle {0082}; Bjorke {0082}; Portal {0082}; Reno {0082}; Tjalve {0082}. Pinb-D1c is a "loss-of-function" mutation resulting from the replacement of a

Pinb-D1d

i	PI 644083 (Alpowa/Mjolner//7*Alpowa) {10429}.
Varieties	Bercy {0082}; Mjolner {0082}; Soissons (homonym 'b1') {10433}. Pinb-D1d is a "loss-of-function" mutation resulting from the replacement of a tryptophan by an arginine

Pinb-D1e

i	PI 644084 (Alpowa/Canadian Red//7*Alpowa) {10429}.
Varieties	Gehun {0204}; Canadian Red {0204}; Chiefkan {0204}; Yunxianxiaomai {10427}. Pinb-D1e is a "loss-of-function" mutation resulting from the replacement of a tryptophan by a

Pinb-D1f

i	PI 644085 (Alpowa/Sevier//7*Alpowa) {10429}.
Varieties	Abyssinia AV12.4 {10430}; The hard component of Utac{0204}. Pinb-D1f is a "loss-of-function" mutation resulting from the replacement of a tryptophan by a stop codon at

Pinb-D1g

i	PI 644086 (Alpowa/Andrews//7*Alpowa) {10429}.
Varieties	Andrews {0204}. Pinb-D1g is a "loss-of-function" mutation resulting from the replacement of a cysteine by a stop codon at position 56 {0204}.

Pinb-D1h

dv	Ae. tauschii TA2369 (GenBank AY251983) Pina-D1c, Gsp-D1c {03105}; TA2527 (GenBank AY251965) Pina-D1c, Gsp-D1e {03105}; TA1649 (GenBank AY251963) PinaD1d, Gsp-D1f {03105}; TA10 (GenBank AY649748)

Pinb-D1i

dv	Ae. tauschii TA2475 (GenBank AY251989) Pina-D1a, Gsp-D1b {03105}; TA2536 (GenBank AY251993) Pina-D1c, Gsp-D1d {03105}; TA2374 (GenBank AY251948) PinaD1d, Gsp-D1d {03105}; TA2512 (GenBank AY251992)

Pinb-D1j

dv	Ae. tauschii TA1599 (GenBank AY251962) Pina-D1a, Gsp-D1g {03105}; TA1691 (GenBank AY251964) Pina-D1a, Gsp-D1h {03105}; Ae. tauschii TA1691 (GenBank AY251946) {03108}.

Pinb-D1k

dv	Ae. tauschii PI554318 (GenBank AJ302108) Pina-D1d {03104}.

Pinb-D1l

Varieties

GaoCheng8901 {10119}. {10208} reported Pinb-D1b in Gaocheng 8901.

Pinb-D1m

Varieties

X. aegilotriticum CIGM87.2783-1B-0PR-0B (GenBank AY573901) Pina-D1c {10118}.

Pinb-D1n

Varieties

X. aegilotriticum CIGM92.1708 (GenBank AY573902) Pina-D1d {10118}.

Pinb-D1o

Varieties

X. aegilotriticum CIGM93.247 (GenBank AY573903) Pina-D1e {10118}.

Pinb-D1p

Synonym	Pinb-D1z {10316}
Varieties	Dahuangpi (GenBank AY581889) {10316}; Nongda 3213 {10121}; Nongda 3395 {10121}; Qindao landrace {10305}; Qitoubai {10305}; Shijiazhuang 34 {10305}; Zigan

Pinb-D1q

Sources / synonyms	CS/Red Egyptian 5D substitution line, Pina-D1k, Gsp-D1i* {10077}.
Varieties	Jingdong 11 (GenBank EF620909) {10313}. This allele was used originally (2004 supplement) in combination with Pina-D1k and Gsp-D1i to denote the large deletion that encompasses

Pinb-D1r

Synonym	Pinb-D1h {10209}
Varieties	Hyb65 (NCBI AJ619022) {10209}. G insertion: open reading frame shift and premature stop codon; hard kernel texture {10209}.

Pinb-D1s

Varieties

NI5439 (NCBI AJ619021) {10209}. G insertion as in Pinb-D1r and an A-to-G substitution; hard kernel texture {10209}.

Pinb-D1t

Varieties

Guangtouxianmai (GenBank EF620910) {10208}; Hongma{10208}. G-to-C substitution: Glycine-47 to arginine; hard kernel texture {10208}

Pinb-D1u

Varieties

Tiekemai (GenBank EF620911) {10427}; 31 hard Yunnan endemic wheats ( T. aestivum ssp. yunnanense King) {10427}. Possesses a G deletion at position 127 leading to

Pinb-D1v

Synonym	Pinb-D1i(t) {10305}, Pinb-D1r {10316}
Varieties	Qingdao Landrace 1 {10305}; Qitoubai {10305}; Shijiazhuang 34 {10305}; Tachun 3 (GenBank AY598029) {10316}; Zigan {10305}; homonym 'b5' {10316}.

The original assignment of this allele in the 2006 supplement was incorrect; the sequence/varieties in {10305] are Pinb-D1p as listed above for that allele. The following variety/sequence was assigned PinbD1y in the 2006 supplement; but the original assignment of {10316} is now unchanged.

Pinb-D1w

Synonym	Pinb-D1q {10316}
Varieties	Jing 771 (GenBank AY640304, AB180737) {10316}; homonym 'b4' {10316}.
dv	Ae. tauschii 002 (GenBank DQ257553) {10314}; Ae. tauschii ssp. tauschii TA1704 (GenBank AY649747) {10315}; Ae. tauschii ssp. anathera TA2381 (GenBank AY649747 {10315}. This variety/sequence was

Ae. tauschii isolate Q03-002 (GenBank DQ257553) (referred to as allele 2) {10314}; Ae. tauschii TA1704 and TA2381 (GenBank AY649747) {10315}; Ae. tauschii CPI 110799 (GenBank CR626926) {10440} were incorrectly assigned this allele in the 2006 supplement; they are Pinb-D1i as listed above.

Pinb-D1x

Varieties

Kashibaipi (GenBank AM909618) {10528}.

Pinb-D1y

The original assignment of this allele in the 2006 supplement was incorrect; the sequence for Tachun 3 in {10305} is Pinb-D1v as listed above. The original assignment of {10316} is now unchanged. Currently there is no assignment for this allele.

Pinb-D1z

This allele/sequence is identical to, and listed under, Pinb-D1p . Currently there is no assignment for this allele.

Pinb-D1aa

Varieties

Changmangtoulongbai (GenBank EF620912) {10391}; Hongtutou 1 {10391}; Hongtutou 2 {10391}.

Pinb-D1ab

Varieties

KU3062 {10432}; KU3069 {10432}; Tuokexunyihao {10528}.

Pinb-D1ac

Varieties

Kashibaipi {10570}; Red Star {10570}.

G to A substitution at position 257 and C to T substitution at position 382 {10570}.

PINa-S1

dv	Ae. speltoides PI 393494 (GenBank AJ302096) {03104}; PI 369616 (GenBank AJ302097) {03104}; Ae. speltoides spp. speltoides TA2368 (GenBank AY622787), TA1789 (GenBank AY622788) {03108}; Ae. speltoides

PINa-S[b] 1

dv	Ae. bicornis spp. typica TA1954, TA1942 {03108}.

Pina-S[l] 1

dv	Ae. longissima spp. longissima TA1912 (GenBank AY622790) {3108}; Ae. longissima spp. nova TA1921 (GenBank AY622791) {03108}.

Pina-S[s] 1

dv	Ae. searsii TA1837, TA1355 (GenBank AY622792) {03108}.

Pina-S[sh] 1

dv	Ae. sharonensis TA1999 (GenBank AY622796) {03108}.

Pinb-D1b, Pinb-D1c, Pinb-D1d, Pinb-D1e, Pinb-D1f , or Pinb-D1g are present in hard hexaploid wheats not carrying the Pina-D1b (null) mutation {452, 1035, 0082, 0204}. Wheats with Pinb-D1b were slightly softer and a little superior to those with Pina-D1b in milling and bread-making characteristics although there was considerable overlap {0206}. Transgenic rice with the Pina-D1a and Pinb-D1a alleles possessed softer grain {0207}. Genotypes for a selection of North American wheats are given in {0204}. In T. monococcum the gene order was reported to be: tel - GSP-1 - PINa - PINb {0083, 10122} whereas in Ae. squarrosa it was: tel - GSP-1 - PINb- PINa {10037}. The soft kernel trait was transferred to durum {10899}. The soft kernel trait was transferred to durum; firstly, to Langdon durum Selection 1-674 and then by backcrossing to cv. Svevo {10899}, which was in turn used to develop backcross derivatives in cv. Alzada, Havasu, Kyle, and Strongfield {11444}. Genetic evidence indicated that ~24.4 Mbp from CS chromosome 5DS replaced ~20 Mbp of 5BS {11444}. Further cytogenetic analysis identified the translocation breakpoint in a 39 bp region within a putative glcosyltransferase gene {11489}. Ikeda et al. {10305} reported a double-null with apparently no PINa-D1 or PINb-D1 genes present in

GLO-1 GLO-A1

Chromosome	1AS
Varieties	CS.
Marker associations	Distally located: GLO-A1 (distal) – 5.2 cM – GLI-A1 {1077}.

GLO-B1

Chromosome	1BS
Varieties	CS.

GLO-D1

Chromosome	1DS
Varieties	CS.
Marker associations	Distally located: GLO-D1 (distal) – 2.9 cM – GLI-D1 {1077}.

GLO-E1

Chromosome	1ES
ad	CS/ E. elongata .

GLO-R1

Chromosome	1RS
ad	CS/Imperial.
su	1B/(1R), eg., Salzmunde 14/44.

SRP-1 SRP-A1

Chromosome

5AL

Srp-B1a

Synonym	Srp5Ba {10754}
Varieties	Etawah {10755}; Federation {10755}; Frame {10755}; Pugsley {10754}; Stylet {10755}.

Srp-B1b

Note	Null allele.
Varieties	Correll {10755}; EGA Eagle Rock {10755}; Gladius {10755}; Yitpi {10755}. This allele reduced milling yield by 0.4% {10755}.

SRP-B1

Synonym	Srp5B {10754}
Chromosome	5BL

SRP-D1

Chromosome

5DL

SGP-2 SGP-A2

Varieties

CS.

Sgp-B2

Varieties

CS.

Sgp-D2

Varieties

CS. .

SGP-3 See also starch synthase,

SSI-1 Sgp-A3

Chromosome	7AS
Varieties	CS.

Sgp-A3a

Varieties

CS.

Sgp-A3b

Note	Null allele.
Varieties	Norin 61.

Sgp-B3

Chromosome	7BS
Varieties	CS.

Sgp-B3a

Varieties

CS.

Sgp-B3b

Note	Null allele.
Varieties	Crest.

Sgp-B3c

Varieties

Spica.

SGP-D3

Chromosome	7DS
Varieties	CS.

A triple null stock (SGP-1 null wheat) is reported in {0137}. Deletion mapping indicated that the gene order on the 7S arms is: centromere - SGP-1 – SGP-3 – Wx {1615}.

SSI-A1

Chromosome

SSI-B1

Chromosome

SSI-D1

Chromosome

SSII-1 . Starch synthase II proteins are identical to the starch granule proteins SGP-1 {0042}

SsII-A1

Chromosome

SsII-B1

Chromosome

SsII-D1

Chromosome

WSP-D1

Chromosome	7DL
Varieties	CS

Wsp-D1a

Varieties

CS.

Wsp-D1b

Varieties

Sears' Synthetic IPSR 1190903.

Wsp-D1c

Varieties

T4 = Agatha {890,893}; Indis {890,892}.

WSP-E1

Chromosome	7E
ad	CS/ E. elongata .

WSP-H1

Chromosome	7H
ad	CS/Betzes.

WSP-H[ch] 1

Chromosome	7H
ad	CS/ H. chilense .

WSP-S[l] 1

Chromosome	7S
ad	CS/ Ae. sharonensis .

WSP-V1

Chromosome	7V
ad	CS/ D. villosum .

Wx-A1a

Synonym	Wx-B1a {1054}
Varieties	Bao Hua {10989}; CS; Hoshuu.
Type varieties	Langdon {10989}.

Wx-A1b

Synonym

Wx-B1b {1054}

Null allele.

Wx-A1c

Varieties

Pakistan Zairaishi selection {10629}; QT105 {1617}; WB6 {1617}.

Wx-A1d

Type varieties

T. dicoccoides KU 8937B {1616}.

Wx-A1e

Type varieties

KU 3659 {10629}; T. durum KU 3655 and KU 3659 {1616}.

Wx-A1f

Note	Null allele.
Varieties	Turkey-124 {10187}; Turkey-140 {10187}; Turkey-171 {10187}; Turkey-280 {10187}; Turkey-299 {10187}. Lines with this allele produce a PCR product with a 173 bp insertion

Wx-A1g

Note	Wx-A1' {10587}.
Varieties	PI 348476 {10587}; Spelt accessions PI 348576 {10587}; 2778 Epeautre Noir Velu {10587}.

Wx-A1h

Note	Null allele.
Type varieties	Buck Topacio {10763}. This is probably a unique allele possessing a 1 bp deletion in exon 6 leading to frameshift and a stop codon: partial

Wx-A1i

Varieties

KU9259{10989}.

Wx-A1j

Varieties

M1 {10989}. Functional markers for Wx-A1c, Wx-A1d, Wx-A1e and Wx-Ali were developed from DNA sequences {10990}.

WX-B1

Synonym	Wx-A1 {1054, 1053}, XWx-4B {179, 180}, XWx-4A {961}
Chromosome	4AL
Varieties	CS.
Type varieties	A variant allele was present in three accessions {03101}. A dominant PCR marker for identifying heterozygotes at the Wx-B1 locus is reported in {10732}.

Wx-B1a

Synonym	Wx-A1a {1054}
Varieties	CS; Joshuu. The complete genomic sequence for Wx-B1a from CS was determined {0073}.

Wx-B1b

Synonym

Wx-A1b {1054}

Null allele.

Wx-B1c

Varieties

AF24 {10629}; Chousen 40 {0094}; Cikotaba {1617, 10629}; Junguk 12 {1617, 10629}.

Wx-B1d

Type varieties

T. durum KU 4213D {1616}; KU 4213D {10629}; KU 4224C {1616}.

Wx-B1e

Varieties

Blue Boy II {0027}; Canthatch {0027}; Eureka {0027}; Gotz {0027}; Norin 44 {0027}; Turkey Red {0027}.

Wx-B1f

Type varieties

BG-12413 {0111}; BG-12415 {0111}.

Wx-B[S] 1g

al	Ae. speltoides 33 {10587}.

Wx-B[SL] 1h

al	Ae. longissima 12 {10587}.

WX-D1 {1053}, {180}. [ XWx-7D {179, 180}]. 7DS {1053}, {180}.

Wx-D1a

Varieties

CS.

Wx-D1b

Note	Null allele.
Varieties	Bai Huo (Baihuomai) {1617}; DHWx12 {0117}.
Varieties (alt.)	Mochi-Otome Wx-A1b Wx-B1b {10032}.
Marker associations	STS marker Xsun1-7D produces a distinct band of about 260 bp (compared with the standard 840 bp), indicative of a smaller PCR product, but the

Wx-D1c

Varieties

Scoutland {1617}.

Wx-D1d

Varieties

K107Wx1 {0118}; K107Wx2 {0118}; One Iranian and one Italian accession {03101}.

Wx-D1e

Note	Null allele {0117}.
Varieties	NP150 {0117}. STS marker Xsun1-7D failed to produce a PCR product {0117

Wx-D1f

Synonym	Wx-d1e {0234}
Varieties	Tanikei A6599-4 {0234}.Relative to Kanto 107, Tanikei A6599-4 carries an alanine to threonine substitution at position 258 in the mature protein {0234}.

Wx-D[DN] 1g

al	Ae. ventricosa 12 {10587}.

Various hard and soft wheats with alleles Wx-A1b, Wx-B1b and Wx-D1b are listed in {0304}. 15% of Chinese wheats possessed Wx-B1 null alleles {10357}. Isolation of genomic sequences for the genes encoding granule-bound starch synthase ( GBSSI or WX ) in T. monococcum, Ae. speltoides and Ae. tauschii was reported in {0168}. Cloning of a second set of GBSSI or waxy genes, GBSSII , which were shown to be located on chromosomes 2AL, 2B and 2D, was reported in {0167}.

GLU-2

Glu-B2a

Note	12 {00114}.
Type varieties	Mexicali.

Glu-B2b

Note	Null {00114}.
Type varieties	Langdon. GLI-B3 was designated GLU-B2 {589} until the name of the locus was changed in {1119}.

Glu-B2c

Note	12* {10215}.
Type varieties	Alcala la Real {10215}.

Glu-B2d

Note	12.1 {11493}.
Type varieties	T. turgidum ssp. durum Moroccan landrace MGB-3125 {11493}.

GLU-3

Glu-A3

Chromosome	1AS
Varieties	CS. The first 7 alleles were distinguished using 5 allele-specific primer sets {10185}. Further mainly Australian genotypes with alleles a to f are listed in

Glu-A3a

Varieties

CS.

Glu-A3b

Varieties

Gabo.

Glu-A3c

Varieties

Cheyenne.

Glu-A3d

Varieties

Cappelle Desprez, Orca; Suneca{10185}.

Glu-A3e

Varieties

Halberd {10185}; Hope, Insignia.

Glu-A3f

Varieties

Rescue.

Glu-A3g

Varieties

Glenlea {10185}.

Glu-A3h

Synonym

Glu-A3d' {03116}

Null {00114}.

Glu-A3i

Note	8[*] +11 {02110}.
Type varieties	Mourisco Fino.

Glu-A3j

Synonym

Glu-A3a {00114}

6 {00114}.

Glu-A3k

Synonym

Glu-A3b {00114}

5 {00114}.

Glu-A3l

Synonym

{00114}

6+10 {00114}.

Glu-A3m

Synonym

Glu-A3d {00114}

6+11 {00114}.

Glu-A3n

Synonym

Glu-A3e {00114}

11 {00114}.

Glu-A3o

Synonym

Glu-A3f {00114}

6+11+20{114}.

Glu-A3p

Synonym

Glu-A3h {00114}

Null {00114}.

Glu-A3q

Synonym

Glu-A3i {10215}

5+20 {10215}.

Glu-A3r

Synonym	Glu-A3d' {03116}
Varieties	Magistral hexaploid triticale {03116}.

Glu-A3s

Synonym

Glu-A3g {00114}

6+10+20 {00114}.

Glu-A3t

Synonym	Glu-A3[m] a {10805}
dv	PI 190947, T. monococcum ssp. monococcum {10805}.

Glu-A3u

Synonym	Glu-A3[m] b {10805}
dv	PI 190946, T. monococcum ssp. monococcum {10805}.

Glu-A3v

Synonym	Glu-A3[m] c {10805}
dv	BGE-020466, T. monococcum ssp. monococcum {10805}.

Glu-A3w

Synonym	Glu-A3[m] d {10805}
dv	PI 191097, T. monococcum ssp. monococcum {10805}.

Glu-A3x

Synonym	Glu-A3[m] e {10805}
dv	BGE-013624, T. monococcum ssp. Monococcum {10805}.

Glu-A3y

Synonym	Glu-A3[m] f {10805}
dv	PI 191094, T. monococcum ssp. monococcum {10805}.

Glu-A3z

Synonym	Glu-A[u] 3-I {10806}
dv	PI 428139, T. urartu {10806}.

Glu-A3aa

Synonym	Glu-A[u] 3-II {10806}
dv	PI 428327, T. urartu {10806}.

Glu-A3ab

Synonym	Glu-A[u] 3-III {10806}
dv	PI 428340, T. urartu {10806}.

Glu-A3ac

Synonym	Glu-A[u] 3-IV {10806}
dv	PI 428322, T. urartu {10806}.

Glu-A3ad

Synonym	Glu-A[u] 3-V {10806}
dv	PI 428188, T. urartu {10806}.

Glu-A3ae

Synonym	Glu-A[u] 3- VI {10806}
dv	PI 428203, T. urartu {10806}.

Glu-A3af

Synonym	Glu-A[u] 3-VII {10806}
dv	PI 428255, T. urartu {10806}.

Glu-A3ag

Synonym	Glu-A[u] 3-VIII {10806}
dv	PI 428328, T. urartu {10806}.

Glu-A3ah

Synonym	Glu-A[u] 3-IX {10806}
dv	PI 428256, T. urartu {10806}.

Glu-A3ai

Synonym	Glu-A[u] 3-X {10806}
dv	PI 428217, T. urartu {10806}.

Glu-A3aj

Synonym	Glu-A[u] 3-XI {10806}
dv	PI 428335, T. urartu {10806}.

Glu-A3ak

Synonym	Glu-A[u] 3-XII {10806}
dv	PI 428186, T. urartu {10806}.

Glu-A3al

Synonym	Glu-A[u] 3-XIII {10806}
dv	PI 428183, T. urartu {10806}.

Glu-A3am

Synonym	Glu-A[u] 3-XIV {10806}
dv	TRI 11563, T. urartu {10806}.

Glu-A3an

Synonym	Glu-A[u] 3-XV {10806}
dv	PI 427328, T. urartu {10806}.

Glu-A3ao

Synonym	Glu-A[u] 3-XVI {10806}
dv	PI 428253, T. urartu {10806}.

Glu-A3ap

Synonym	Glu-A[u] 3-XVII {10806}
dv	PI 538735, T. urartu {10806}.

Glu-A3aq

Synonym	Glu-A[u] 3-XVIII {10806}
dv	PI 428225, T. urartu {10806}.

Glu-A3ar

Synonym	Glu-A[u] 3-XIX {10806}
dv	PI 538733, T. urartu {10806}.

Glu-A3as

Synonym	Glu-A[u] 3-XX {10806}
dv	PI 428196, T. urartu {10806}.

Glu-A3at

Synonym	Glu-A[u] 3-XXI {10806}
dv	PI 538724, T. urartu {10806}.

Glu-A3au

Synonym	Glu-A[u] 3-XXII {10806}
dv	PI 428191, T. urartu {10806}.

Glu-A3av

Synonym	Glu-A[u] 3-XXIII {10806}
dv	TRI 6734, T. urartu {10806}.

Glu-A3aw

Synonym	Glu-A[u] 3-XXIV {10806}
dv	TRI 11496, T. urartu {10806}.

Glu-A3ax

Note	6.1 {10116}.
Type varieties	Buck Cristal {10116}. The designation of this protein (subunit 6.1) as encoded by GLU-A3 , previously deduced from its electrophoretic mobility {10116}, was confirmed through

Glu-A3ay

Note	6+20 {11492}.
Type varieties	T. turgidum ssp. durum landraces BGE047515 and BGE047516 {11492}; Mexican durum landrace accession 10 (CWI52016) {11490}.

Glu-A3az

Note	6+10+11* {11490}.
Type varieties	T. turgidum ssp. durum Mexican landrace accession 3 (CWI51941) {11490}.

Glu-A3ba

Note	5+11 {11492}.
Type varieties	T. turgidum ssp. turgidum landrace BGE047535 {11492}; Iranian landrace accession 77 (CWI73342) {11490}.

Glu-A3bb

Note	20 {11492}.
Type varieties	T. turgidum ssp. dicoccon landrace BGE047498 {11492}; T. turgidum ssp. turgidum landrace BGE047531 {11492}; T. turgidum ssp. durum Moroccan landrace MGB-16563 {11493}.

Glu-A3bc

Note	5** {11493}.
Type varieties	T. turgidum ssp. durum Moroccan cv. MGB-20 {11493}. Glu-A3bd [{11492}]. 5+22 {11492}.
Type varieties	T. turgidum ssp. turgidum BGE047532 {11492}. Glu-A3be [{11492}]. 5* {11492}.
Type varieties	T. turgidum ssp. turgidum BGE048495 {11492}. Glu-A3bf [{11492}]. 5+20 {11492}. tv : T. turgidum* ssp. turgidum BGE048498 {11492}. Glu-A3bg [{11539}]. 5*+11+20 {11539}.
Type varieties	T. turgidum ssp. turgidum BGE018646 {11539}. Glu-A3bh [{11539}]. 10 {11539}.
Type varieties	T. turgidum ssp. durum BGE013622 {11539}. Glu-A3bi [{11539}]. 5*+11+22 {11539}.
Type varieties	T. turgidum ssp. turgidum BGE013089 {11539}. Glu-A3bj [{11540}]. 5* {11540}.
Type varieties	T. turgidum ssp. durum Fanfarron {11540}. Glu-A3bk [{11540}]. 8* {11540}.
Type varieties	T. turgidum ssp. durum BGE019300 {11540}. Glu-A3bl [{11540}]. 5+8* {11540}.
Type varieties	T. turgidum ssp. durum BGE013718 {11540}.

GLU-B3

Chromosome	1BS
Varieties	CS. Three different approaches were employed to identify putative SNPs used to design gene-specific primers for LMW-GS genes, and six functional STS markers, three for

Glu-B3a

Varieties

CS.

Glu-B3b

Varieties

Gabo, Timstein, Hope.

Glu-B3c

Varieties

Insignia, Halberd.

Glu-B3d

Varieties

Orca.

Glu-B3e

Varieties

Cheyenne.

Glu-B3f

Varieties

Radja.

Glu-B3g

Varieties

Kharkov, Bungulla.

Glu-B3h

Varieties

Thatcher, Rescue.

Glu-B3i

Varieties

Norin-61.

Glu-B3j

Note	.
Varieties	Kavkaz. Null allele carried by the 1BL.1RS translocation. This allelic designation was originally incorrectly used in the catalogue to name an allele from T. turgidum

Glu-B3k

Note	8+9+13+16+19 {02110}.
Type varieties	ALP-153, Dural, Durati, Edmore; Faisca.

Glu-B3l

Type varieties

Gionp-1954.

Glu-B3m

Synonym	Glu-B3b' {03120}
Varieties	Soissons {03120}.

Glu-B3n

Synonym	Glu-B3c' {03120}
Varieties	Courtot {03120}.

Glu-B3o

Synonym	Glu-B3i' {03116}
Varieties	Olympus hexaploid triticale {03116}.

Glu-B3p

Synonym	Glu-B3k {03116}
Varieties	Alamo hexaploid triticale {03116}.

Glu-B3q

Synonym	Glu-B3h' {03115}
Varieties	Torote hexaploid triticale {03115}

Glu-B3r

Synonym

Glu-B3a {00114}

2+4+15+19 {00114}.

Glu-B3s

Synonym

Glu-B3b {00114}

8+9+13+16 {00114}.

Glu-B3t

Synonym

Glu-B3c {00114}

2+4+14+15+19 {00114}.

Glu-B3u

Synonym

Glu-B3d {00114}

2+4+15+17+19 {00114}.

Glu-B3v

Synonym

Glu-B3e {00114}

2+4+15+16+18 {00114}.

Glu-B3w

Synonym

Glu-B3f {00114}

2+4+15+17 {00114}.

Glu-B3x

Synonym

Glu-B3g {00114}

2+4+15+16 {00114}.

Glu-B3y

Synonym

Glu-B3h {00114}

1+3+14+18 {00114}.

Glu-B3z

Note	6.1 {10116}.
Type varieties	Buck Cristal {10116}.

Glu-B3aa

Synonym

Glu-B3l {10215}

1+3+13*+16 {10215}.

Glu-B3ab

Varieties

Hope {10804}; Nanbukomugi {10804}.

Glu-B3ac

Varieties

ACA 801 {10804}; Klein Proteo {10804}; Thesee {10804}.

Glu-B3ad

Varieties

AC Vista {10804}; Heilo {10804}; Opata 85 {10804}; Ruso {10804}.

Glu-B3ae

Note	1+3+16 {11490}.
Type varieties	T. turgidum ssp. durum accession 56 (CWI57386) {11490}.

Glu-B3af

Note	1+3+17 {11490}.
Type varieties	T. turgidum ssp. durum accession 74 (CWI71827) {11490}.

Glu-B3ag

Note	2+4+16 {11490}.
Type varieties	T. turgidum ssp. durum accession 46 (CWI56913) {11490}.

Glu-B3ah

Note	8+9+16 {11490}.
Type varieties	T. turgidum ssp. durum accession 65 (CWI57719) {11490}.

Glu-B3ai

Note	2+4+14+18 {11490}.
Type varieties	T. turgidum ssp. durum accession 62 (CWI57615) {11490}.

Glu-B3aj

Note	19 {11490}.
Type varieties	T. turgidum ssp. durum accession 77 (CWI73342) {11490}.

Glu-B3ak

Note	2+4+6*+15+19 {11490}.
Type varieties	T. turgidum ssp. durum accession 69 (CWI71627) {11490}.

Glu-B3al

Note	2+4+7*+15+16 {11490}.
Type varieties	T. turgidum ssp. durum accession 61 (CWI57614) {11490}.

Glu-B3am

Note	1+3+6*+13’+17 {11490}.
Type varieties	T. turgidum ssp. durum accession 72 (CWI71759) {11490}.

Glu-B3an

Note	8+9+13’+17 {11490}.
Type varieties	T. turgidum ssp. durum accession 58 (CWI57522) {11490}.

Glu-B3ao

Note	1+3+8’+17 {11490}.
Type varieties	T. turgidum ssp. durum accession 50 (CWI57256) {11490}.

Glu-B3ap

Note	2+4+6*+9’+14+19 {11490}.
Type varieties	T. turgidum ssp. durum accession 78 (CWI73350) {11490}.

Glu-B3aq

Note	2+4+8+15+19 {11492}.
Type varieties	T. turgidum ssp. durum landraces BGE045634 {11492}; MGB-2963 {11493}.

Glu-B3ar

Note	1+3+7+15+18 {11493}.
Type varieties	T. turgidum ssp. durum landrace MGB-16563 {11493}.

Glu-B3as

Note	1+3+8+13+16+19 {11493}.
Type varieties	T. turgidum ssp. durum landrace MGB-3152 {11493}.

Glu-B3at

Note	2+4+17 {11493}.
Type varieties	T. turgidum ssp. durum landrace MGB-3125 {11493}.

Glu-B3au

Note	2+4+7+15+19 {11493}.
Type varieties	T. turgidum ssp. durum landrace MGB-5963 {11493}.

Glu-B3av

Note	9+13+16 {11493}.
Type varieties	T. turgidum ssp. durum landrace MGB-3101 {11493}. Glu-B3aw [{11492}]. 1+3+8+13+15+18 {11492}.
Type varieties	T. turgidum ssp. turgidum BGE047502 {11492}. Glu-B3ax [{11492}]. 1+3+13*+19 {11492}.
Type varieties	T. turgidum ssp. turgidum BGE047504, BGE047506 {11492}. Glu-B3ay [{11492}]. 1+3+14+15 {11492}.
Type varieties	T. turgidum ssp. turgidum BGE047521 {11492}. Glu-B3az [{11492}]. 1+16 {11492}.
Type varieties	T. turgidum ssp. dicoccon BGE045645, BGE047503 {11492}. Glu-B3ba [{11492}]. 2+4+7+13*+15+19 {11492}.
Type varieties	T. turgidum ssp. durum BGE045651 {11492}. Glu-B3bb [{11492}]. 2+4+15 {11492}.
Type varieties	T. turgidum ssp. durum BGE047516 {11492}. Glu-B3bc [{11492}]. 2+4+15+17+21 {11492}.
Type varieties	T. turgidum ssp. turgidum BGE048494 {11492}. Glu-B3bd [{11492}]. 4+(7**)+13+15+19 {11492}.
Type varieties	T. turgidum ssp. dicoccon BGE045628 {11492}.

The designation of subunit 7 as encoded by _Glu-A3_ was deduced from its electrophoretic mobility and awaits confirmation through mapping studies {11492}; the subunit was therefore referenced by {11492}. _Glu-B3be_ [{11492}]. 4+(7)+13+15+21 {11492}.

Glu-D3a

Varieties

CS.

Glu-D3b

Varieties

Gabo.

Glu-D3c

Varieties

Insignia, Cappelle Desprez.

Glu-D3d

Varieties

Jufy-1 {10813}; Norin-61A.

Glu-D3e

Varieties

Orca, Thatcher.

Glu-D3f

Varieties

Cheyenne {10548}.

Glu-D3g

Varieties

Hira-1 {10558}.

Glu-D3h

Varieties

India 115 {10558}.

Glu-D3i

Varieties

Bolac {10558}.

Glu-D3j

Varieties

Hira-2 {10558}.

Glu-D3k

Varieties

Lincoln {10558}.

Glu-D3l

Varieties

Heilo {10804}; Jing411 {10804}; Pepital {10804}; Thesee {10804}.

Glu-D3m

Varieties

Darius {10804}.

Glu-D3n

Varieties

Fengmai 27 {10804}.

GLU-E3

Chromosome	1ES
su	CS/ E. elongata .

GLU-S[l] 3 {1228}, {480}. 1S[l] {480}. 1S[1] S {1228}.

Glu-U3

Chromosome	1U
su	CS/ Ae. umbellulata .

A series of papers {00106, 00107, 00108 and 00109} describe considerable variation in primitive wheats not present in bread wheat (A genome species T. boeoticum, T. urartu, T. thaoudar, T. aegilopoides, T. monococcum , and D-genome species T. tauschii ) for the low molecular weight subunits, sufficient to use them as a source for potentially changing flour properties in bread wheat. In {00110}, variants for LMW glutenin subunits were reported from study of 24 accessions of einkorn wheat ( T. monococcum ssp. monococcum ). Nine of these showed two electrophoretic bands for LMW subunits, arbitrarily designated 'a' and 'b', that appeared to be associated with good bread-making quality. The isolation of a new low-molecular-weight glutenin subunit gene, located on chromosome 1D, was reported in {0350}.

GLU-4

Glu-D4

Chromosome	1D
su	CS/Langdon 1D(1A); CS/Langdon 1D(1B) {02111}.

Glu-D4a

Varieties

J 24.

Glu-D4b

Varieties

PBW 154.

Glu-D4c

Note	Null allele.
Varieties	NI 4.

GLU-5

GLU-D5

Chromosome	7D
su	CS/Langdon 7D(7A); CS/Langdon 7D(7B) {02111}.

A collection of 173 Ae. tauschii accessions were analysed for low molecular weight glutenin subunits by SDS-PAGE {02112}; 33 different patterns for B-subunits and 43 for C-subunits were identified, some of which were of identical electrophoretic mobility to those observed in common wheat. Also observed were subunits with the same mobilities as the D-subunits and as the subunits encoded by the GLU-D4 and GLU-D5 loci. This variation represents a source of novel germplasm of potential value for breeding programmes aimed at improving the D-genome of common wheat in the context of bread-making quality.

Glu-D5a

Varieties

PBW 154.

Glu-D5b

Note	Null allele.
Varieties	K 68.

GLI-1

GLI-A1

Synonym

Gld 1A {1415}

1AS {150}, {634}, {1607}, {1334}.

Gli-A1a

Varieties

Castan {991}; CS {988}; Mara {9986}; Mentana {9986}; Millewa {00119}.

Gli-A1b

Varieties

Bezostaya 1, Mercia {988}; Tracy {991}.

Gli-A1c

Varieties

Ukrainka {998}; Gazul {9985}; Sava {994}; Hopps {00119}.

Gli-A1d

Varieties

Dankowska {988}; Cabezorro {9985}.

Gli-A1e

Varieties

Falchetto {988}; Open {991}; Touzelle {991}.

Gli-A1f

Varieties

Mironovskaya 808, Maris Freeman {988}; Arminda {991}. Note: An allele Gli-A1f * is mentioned in {03130}.

Gli-A1g

Varieties

Gabo {988}; Adalid {9985}.

Gli-A1h

Varieties

Sadovo I {988}; Predela {9981}; Krajinka {9981}.

Gli-A1i

Varieties

Saratovskaya 36 {988}.

Gli-A1j

Varieties

Lutescens 62 {988}.

Gli-A1k

Varieties

Courtot {991}; Skala (heterogeneous) {988}; Soissons {991}; Spada {9986}.

Gli-A1l

Varieties

Lesostepka 75 {988}; David {9986}; Salmone {9986}; Mura {9981}.

Gli-A1m

Varieties

Marquis {988}; Dneprovskaya 521 {988}; Carat {991}; Liocorno {9986}.

Gli-A1n

Varieties

Intensivnaya {988}.

Gli-A1o

Varieties

Odesskaya 16 (heterogeneous) {988}; Oderzo {9986}; Cappelle-Desprez {991}; Capitole {991}.

Gli-A1p

Varieties

Pyrotrix 28 {988}; Zagore {9981}.

Gli-A1q

Varieties

Akmolinka 1 {988}.

Gli-A1r

Varieties

Ranniaya 73 {988}; Barbilla {9985}.

Gli-A1s

Although reported {9986}, this allele is omitted because it requires further confirmation {9981}.

Gli-A1t

Varieties

Jeja del Pais {9985}; Milturum 553 {9981}; Strela {9981}.

Gli-A1u

Varieties

Candeal Alcala {9985}.

Gli-A1v

Varieties

Japhet {9981}; Rouge de Bordeaux {9981}.

Gli-A1w

Note	Null allele.
Varieties	Saratovskaya 29 (mutant) {9987}; E. Mottin {9981}.

Gli-A1x

Synonym	Gli-A1[m] a {10805}
dv	PI 191146, T. monococcum ssp. monococcum {10805}.

Gli-A1y

Synonym	Gli-A1[m] b {10805}
dv	PI 190947 T. monococcum ssp. monococcum {10805}.

Gli-A1z

Synonym	Gli-A1[m] c {10805}
dv	PI 190946, T. monococcum ssp. monococcum {10805}.

Gli-A1aa

Synonym	Gli-A1[m] d {10805}
dv	PI 191097, T. monococcum ssp. monococcum {10805}.

Gli-A1ab

Synonym	Gli-A1[m] e {10805}
dv	BGE-020466, T. monococcum ssp. monococcum {10805}.

Gli-A1ac

Synonym	Gli-A1[m] f {10805}
dv	BGE-013626, T. monococcum ssp. monococcum {10805}.

Gli-A1ad

Synonym	Gli-A1[m] g {10805}
dv	BGE-013628, T. monococcum ssp. monococcum {10805}.

Gli-A1ae

Synonym	Gli-A[u] 1-I {10811}
dv	PI-428333, T. urartu {10811}.

Gli-A1af

Synonym	Gli-A[u] 1-II {10811}
dv	PI-428319, T. urartu {10811}.

Gli-A1ag

Synonym	Gli-A[u] 1-III {10811}
dv	PI-428335, T. urartu {10811}.

Gli-A1ah

Synonym	Gli-A[u] 1-IV {10811}
dv	PI-428323, T. urartu {10811}.

Gli-A1ai

Synonym	Gli-A[u] 1-V {10811}
dv	PI-428231, T. urartu {10811}.

Gli-A1aj

Synonym	Gli-A[u] 1-VI {10811}
dv	PI-428194, T. urartu {10811}.

Gli-A1ak

Synonym	Gli-A[u] 1-VII {10811}
dv	PI-428256, T. urartu {10811}.

Gli-A1al

Synonym	Gli-A[u] 1-VIII {10811}
dv	PI-428234, T. urartu {10811}.

Gli-A1am

Synonym	Gli-A[u] 1-IX {10811}
dv	PI-428320, T. urartu {10811}.

Gli-A1an

Synonym	Gli-A[u] 1-X {10811}
dv	PI-428255, T. urartu {10811}.

Gli-A1ao

Synonym	Gli-A[u] 1-XI {10811}
dv	PI-428241, T. urartu {10811}.

Gli-A1ap

Synonym	Gli-A[u] 1-XII {10811}
dv	PI-428235, T. urartu {10811}.

Gli-A1aq

Synonym	Gli-A[u] 1-XIII {10811}
dv	PI-428183, T. urartu {10811}.

Gli-A1ar

Synonym	Gli-A[u] 1-XIV {10811}
dv	PI-428317, T. urartu {10811}.

Gli-A1as

Synonym	Gli-A[u] 1-XV {10811}
dv	PI-427328, T. urartu {10811}.

Gli-A1at

Synonym	Gli-A[u] 1-XVI {10811}
dv	PI-428327, T. urartu {10811}.

Gli-A1au

Synonym	Gli-A[u] 1-XVII {10811}
dv	PI-428253, T. urartu {10811}.

Gli-A1av

Synonym	Gli-A[u] 1-XVIII {10811}
dv	PI-428224, T. urartu {10811}.

Gli-A1aw

Synonym	Gli-A[u] 1-XIX {10811}
dv	PI-538727, T. urartu {10811}.

Gli-A1ax

Synonym	Gli-A[u] 1-XX {10811}
dv	PI-428211, T. urartu {10811}.

Gli-A1ay

Synonym	Gli-A[u] 1-XXI {10811}
dv	PI-538724, T. urartu {10811}.

Gli-A1az

Synonym	Gli-A[u] 1-XXII {10811}
dv	PI-428191, T. urartu {10811}.

Gli-A1ba

Synonym	Gli-A[u] 1-XXIII {10811}
dv	TRI-6735, T. urartu {10811}.

Gli-A1bb

Synonym	Gli-A[u] 1-XXIV {10811}
dv	TRI-11494, T. urartu {10811}.

Gli-A1bc

Synonym	Gli-A[u] 1-XXV {10811}
dv	TRI-6734, T. urartu {10811}.

Gli-A1bd

Synonym	Gli-A[u] 1-XXVI {10811}
dv	TRI-11496, T. urartu {10811}.

GLI-B1

Synonym	Gld 1B {1415, 1243}, Gld-B4 {420}, Gld-B2 {420}, Gld-B6 {420}, Gld-B5 {420}, Gld-B3 {420}, Gld-B1 {420}
Chromosome	1BS
Chromosome	1B
Sources / synonyms	CS[*] /Cheyenne {634}.
Varieties	CS {150}, {1607}.

Gli-B1a

Varieties

CS {988}.

Gli-B1b

Varieties

Bezostaya 1 {988}; Carat {991}; Marquis {988}; Liocorno {9986}; Soissons {991}.

Gli-B1c

Varieties

Siete Cerros 66 {988}; Prinqual {991}; Loreto {9986}.

Gli-B1d

Varieties

Dneprovskaya 521 {988}; Chopin {991}; Petrel {991}; Tiberio {9986}; Yecora {9985}; Neepawa {995}; Suneca {00119}.

Gli-B1e

Varieties

Apexal {991}; Fournil {991}; Lutescens 62 {988}; Oderzo {9986}.

Gli-B1f

Varieties

Capitole {991}; Cappelle-Desprez {991}; Dankowska {988}; Maris Freeman {988}; Mercia {998}.

Gli-B1g

Varieties

Champtal {991}; Galahad {988}; Mara {9986}; Sadovo 1 {988}; Tracy {991}.

Gli-B1h

Varieties

Cabezorro {9985}; Krasnodonka {988}; Pepital {991}; Rudi {991}; Tincurrin {00119}.

Gli-B1i

Varieties

Ghurka {988}; Insignia {988}.

Gli-B1j

Varieties

Cluj 650 {988}.

Gli-B1k

Varieties

Crverkapa {994}; De Carolis {9986}; Kremena {988}; Mentana {9986}.

Gli-B1l

Varieties

Avrova {9981}; Clement {991}; Damier {991}; Fiocco {9986}; Kavkaz {9981}. Gli-B1l encodes secalins ssociated with the 1BL.1RS translocation.

Gli-B1m

Varieties

Costantino {9986}; Et.d'Choisy {991}; Pyrotrix 28 {988}.

Gli-B1n

Varieties

Intensivnaya {988}.

Gli-B1o

Varieties

Aragon 03 {9985}; Levent {988}; Pippo {9986}; San Rafael {9985}.

Gli-B1p

Varieties

Inia 66 {9985}; New Pusa 834 {988}.

Gli-B1q

Varieties

Gallo {9986}; Goelent {991}; Goya {991}.

Gli-B1r

Varieties

Chinook {995}; Gazul {9985}; Sevillano {9985}.

Gli-B1s

Varieties

Salmone {9986}; Resistente {9986}; E. Mottin {9981}.

Gli-B1t

Varieties

Jeja del Pais {9985}.

Gli-B1u

Varieties

Negrillo {9985}.

Gli-B1v

Varieties

Montjuich {9985}.

Gli-B1w

Varieties

Ardica{9981}; Barbilla (MCB-1017) {9981}.

Gli-B1x

Note	Null allele.
Varieties	Touzelle {991}; Florence Aurora {9985}. Twelve microsatellite alleles plus a null allele were found at the GLI-B1 locus tightly linked to GLU-B3 in 112

GLI-D1 {121}, {1125}. [ Gld-D1 {420}, Gld-D3 {420}, Gld 1D {1415}, Gld-D2 {420}]. 1DS {150}, {121}, {634,} {1607}, {1334}.

Gli-D1a

Varieties

CS {988}; Marquis {988}; Mentana {9986}; Prinqual {991}; Saratovskaya 36 {988}.

Gli-D1b

Varieties

Bezostaya 1 {988}; Cappelle-Desprez {991}; Etoile d'Choisy {991}; Galahad {988}.

Gli-D1c

Varieties

Skorospelka Uluchshennaya (biotype) {988, 9982}.

Gli-D1d

Varieties

De Carolis {9986}; Solo {988}.

Gli-D1e

Varieties

Gerek 79 {988}.

Gli-D1f

Varieties

Carlos {991}; Gabo {988}; Maris Freeman {988}; Orso {9986}.

Gli-D1g

Varieties

Fournil {991}; Ghurka {988}; Mironovskaya 808 {988}; Open {991}.

Gli-D1h

Varieties

Sadovo I {988}; Zlatostrui {9981}.

Gli-D1i

Varieties

Insignia {988}; Napayo (biotype) {995}; San Rafael {9985}; Tselinogradka {988}.

Gli-D1j

Varieties

Aubain; Chinook {995}; Inia 66 {9985}; Petrel {991}; Promin {988}.

Gli-D1k

Varieties

Cargimarec {991}; Kremena {988}; Mara {9986}; Pippo {9986}.

Gli-D1l

Varieties

Artaban {991}; Corin {991}; Longbow {988}.

Gli-D1m

Varieties

Heurtebise {991}.

Gli-D1n

Varieties

Blanquillo de Toledo (MCB-0950) {9981}.

Gli-D1o

Note	Null allele.
Varieties	Darius {991}; Touzelle {991}; Saratovskaya 29 (mutant) {9987}.

GLI-Ag[i] 1

Note	1Ag[i ] {168}.
ad	Vilmorin 27/ Th. intermedium .

GLI-E1

Chromosome	1ES
ad	CS/ E. elongata .

GLI-H[t] 1

Note	1H[t] p {1037}.
ad	CS/ E. trachycaulum .

GLI-R1

Synonym

SecR1 {1356}, Sec1 {1336}

1RS {1340}, {781}, {1336,} {1334}.

GLI-R[m] 1

Note	1R[m] S {1340}.
ad	CS/ S. montanum .

GLI-S[l] 1

Chromosome	1S
ad	CS/ Ae. longissima .

GLI-U1

Note	1U {150}, {1335}.
ad	CS/ Ae. umbellulata . GLI-V1 {111}, {1026}. 1V {111}, {1026}.
ad	CS/ D. Villosum {1026}; Creso/ D. villosum {111}.

In barley, the B and C hordeins are controlled by the HOR2 and HOR1 loci, respectively, which are linked {1341} on chromosome 1HS {1063, 1153}. The map distances and homology of the proteins indicate that HOR1 , the locus closest to the centromere, is equivalent to the omega-gliadins ( GLI-1-1 ) in GLI-1 {1338}. Three alleles at each of the GLI-1-1 (omega gliadin) loci were noted {1358}. The complexity of the GLI1 compound loci is further emphasized by a report of individual genes being separable by recombination, where G1d-1A (a block of gamma and omega genes) is separable by 0.3% from Gld4-1A (omega gliadins) which is in turn, separable by 1.5% from Gld3-1A (omega gliadins) {1103}. Variation was described elsewhere {634}, {996}, {1126} and applied in mapping experiments {107}, {196}, {422}, {1120}, {1125}, {1243}. Sixteen combinations of GLI-B1 and 4 combinations of GLI-D1 subunits are listed in {420}. Multiple alleles described in {996}, number 15 at GLI-A1 , 18 at GLI-B1 , and 8 at GLI-D1 . The Gli-1 alleles present in 57 Yugoslav wheat varieties were reported in {994}.

Gli-DT1

Chromosome	1DS
Varieties	L/18913 (synthetic).
dv	Ae. tauschii AUS18913. A locus designated GLI-DT1 controlling an omega-gliadin of Ae. tauschii was mapped on the short arm of chromosome 1D between loci GLI-D1

Gli-DT1a

Note	T1.
Varieties	L/18913 (synthetic).
dv	Ae. tauschii AUS18913.

A 1,200 bp Dra I RFLP was identified as a gene-specific probe for the T1 omega-gliadin {10645}.

GLI-2

GLI-A2

Synonym	Gld 6A {1415}
Chromosome	6A
Chromosome	6AS
Varieties	CS.

Gli-A2a

Varieties

Cabezorro {9985}; CS {988}; Insignia {988}; Rieti DIV {9986}.

Gli-A2b

Varieties

Aradi {9985}; Bezostaya 1 {988}; Rivoli {991}; Tiberio {9986}.

Gli-A2c

Varieties

Eagle {00119}; Escualo {9985}; Loreto {9986}; Prinqual {991}; Siete Cerros 66 {988}.

Gli-A2d

Varieties

Dneprovskaya 521 {988}; Kenyon (biobype) {995}; Mocho Sobarriba {9985}.

Gli-A2e

Varieties

Cobra {991}; Mentana {9986}; Resistente {9986}; Sadovo 1 {988}; Sevillano {9985}.

Gli-A2f

Varieties

Adalid {9985}; Gala {991}; Maris Freeman {988}; Sistar {9986}.

Gli-A2g

Varieties

Cappelle-Desprez {991}; Ducat {988}; Mahissa 1 {9985}; Mara {9986}.

Gli-A2h

Varieties

Apollo {991}; Basalt {9981}; Hereward {988}; Montjuich {9985}; N. Strampelli {9986}.

Gli-A2i

Varieties

Krasnodonka {988}; Lesostepka 75 {988}.

Gli-A2j

Varieties

Avalon {9981}; Camp Remy {991}; E. Mottin {9981}; Recital {991}.

Gli-A2k

Varieties

Akmolinka 1 {988}; Estica {991}; Pyrotrix 28 {988}; Renan {991}; Zena {9986}.

Gli-A2l

Varieties

Chamorro {9985}; Champlein {991}; Longbow {988}.

Gli-A2m

Varieties

Marquis {988}; Rex {991}; Suneca {00119}.

Gli-A2n

Varieties

Mironovskaya 808 {988}.

Gli-A2o

Varieties

Calatrava {9985}; Castan {991}; Glenwari {9981}; Lontra {9986}; Touzelle {991}.

Gli-A2p

Varieties

Cajeme 71 {9985}; Capitole {991}; Clement{991}; Pliska {988}; S. Lorenzo {9986}; Yecora 70 {9985}.

Gli-A2q

Varieties

Candeal Alcala {9985}; Montcada {9985}; Saratovskaya 39 {988}.

Gli-A2r

Varieties

Genia l {991}; Open{991}; Riband {988}.

Gli-A2s

Varieties

Saratovskaya 36 {998}.

Gli-A2t

Varieties

Courtot {991}; Prostor{9981}; Rinconada {9985}; Soissons {991}.

Gli-A2u

Varieties

Aragon 03 {9985}; Kirgizskaya Yubileinaya {988}; Saunders {995}; Titien {991}.

Gli-A2v

Varieties

Kzyl-Bas {988}.

Gli-A2w

Varieties

Bezenchukskaya 98 (biotype) {988}.

Gli-A2x

Varieties

Solo {988}.

Gli-A2y

Varieties

Gentil Rosso 202 {9981}; PI 191245 {9981}.

Gli-A2z

Varieties

Gallo {9986}; Giuliana {9986}.

Gli-A2aa

Varieties

Navarro 122 {9985}.

Gli-A2ab

Varieties

Navarro 150 {9985}.

Gli-A2ac

Varieties

Blanquillo de Barcarrota (MCB-0893) {9981}.

Gli-A2ad

Varieties

Hembrilla Soria (MCB-1298) {9981}.

Gli-A2ae

Varieties

Candeal de S.Lorenzo Parrilla (MCB-0932) {9981}.

Gli-A2af

Varieties

Barbilla de Leon (MCB-1292) {9981}.

Gli-A2ag

Varieties

Gluclub {9981}; Tincurrin {9981}.

Gli-A2ah

Varieties

Candeal de Nava del Rey (MCB-0892) {9981}.

Gli-A2ai

Varieties

Blanquillo (MCB-0908) {9981}.

Gli-A2aj

Note	Null allele.
Varieties	Saratovskaya 29 (mutant){9987}.

Gli-A2ak

Synonym	Gli-A2[m] a {10805}
dv	BGE-013630, T. monococcum ssp. monococcum {10805}.

Gli-A2al

Synonym	Gli-A2[m] b {10805}
dv	PI 094740, T. monococcum ssp. monococcum {10805}.

Gli-A2am

Synonym	Gli-A2[m] c {10805}
dv	PI 190942, T. monococcum ssp. monococcum {10805}.

Gli-A2an

Synonym	Gli-A2[m] d {10805}
dv	PI 190947, T. monococcum ssp. monococcum {10805}.

Gli-A2ao

Synonym	Gli-A2[m] e {10805}
dv	PI 190946, T. monococcum ssp. monococcum {10805}.

Gli-A2ap

Synonym	Gli-A2[m] f {10805}
dv	BGE-013626, T. monococcum ssp. monococcum {10805}.

Gli-A2aq

Synonym	Gli-A2[m] g {10805}
dv	PI 191095, T. monococcum ssp. monococcum {10805}.

Gli-A2ar

Synonym	Gli-A2[m] h {10805}
dv	BGE-001937, T. monococcum ssp. monococcum {10805}.

Gli-A2as

Synonym	Gli-A2[m] i {10805}
dv	PI 191096, T. monococcum ssp. monococcum {10805}.

Gli-A2at

Synonym	Gli-A2[m] j {10805}
dv	BGE-020466, T. monococcum ssp. monococcum {10805}.

Gli-A2au

Synonym	Gli-A2[m] k {10805}
dv	BGE-001937, T. monococcum ssp. monococcum {10805}.

Gli-A2av

Synonym	Gli-A2[m] l {10805}
dv	BGE-029108, T. monococcum ssp. monococcum {10805}.

Gli-A2aw

Synonym	Gli-A2[m] m {10805}
dv	BGE-013627, T. monococcum ssp. monococcum {10805}.

Gli-A2ax

Synonym	Gli-A2[m] n {10805}
dv	BGE-001937, T. monococcum ssp. monococcum {10805}.

Gli-A2ay

Synonym	Gli-A[u] 2-I {10811}
dv	PI-428333, T. urartu {10811}.

Gli-A2az

Synonym	Gli-A[u] 2-II {10811}
dv	PI-428320, T. urartu {10811}.

Gli-A2ba

Synonym	Gli-A[u] 2-II {10811}
dv	PI-428230, T. urartu {10811}.

Gli-A2bb

Synonym	Gli-A[u] 2-IV {10811}
dv	PI-428319, T. urartu {10811}.

Gli-A2bc

Synonym	Gli-A[u] 2-V {10811}
dv	PI-428239, T. urartu {10811}.

Gli-A2bd

Synonym	Gli-A[u] 2-VI {10811}
dv	PI-428336, T. urartu {10811}.

Gli-A2be

Synonym	Gli-A[u] 2-VII {10811}
dv	PI-428235, T. urartu {10811}.

Gli-A2bf

Synonym	Gli-A[u] 2-VIII {10811}
dv	PI-428234, T. urartu {10811}.

Gli-A2bg

Synonym	Gli-A[u] 2-IX {10811}
dv	PI-428183, T. urartu {10811}.

Gli-A2bh

Synonym	Gli-A[u] 2-X {10811}
dv	PI-428256, T. urartu {10811}.

Gli-A2bi

Synonym	Gli-A[u] 2-XI {10811}
dv	PI-428255, T. urartu {10811}.

Gli-A2bj

Synonym	Gli-A[u] 2-XII {10811}
dv	PI-428224, T. urartu {10811}.

Gli-A2bk

Synonym	Gli-A[u] 2-XIII {10811}
dv	PI-428208, T. urartu {10811}.

Gli-A2bl

Synonym	Gli-A[u] 2-XIV {10811}
dv	PI-428202, T. urartu {10811}. ]

Gli-A2bm

Synonym	Gli-A[u] 2-XV {10811}
dv	PI-428217, T. urartu {10811}.

Gli-A2bn

Synonym	Gli-A[u] 2-XVI {10811}
dv	PI-427328, T. urartu {10811}.

Gli-A2bo

Synonym	Gli-A[u] 2-XVII {10811}
dv	PI-428317, T. urartu {10811}.

Gli-A2bp

Synonym	Gli-A[u] 2-XVIII {10811}
dv	PI-428253, T. urartu {10811}.

Gli-A2bq

Synonym	Gli-A[u] 2-XIX {10811}
dv	PI-538742, T. urartu {10811}.

Gli-A2br

Synonym	Gli-A[u] 2-XX {10811}
dv	PI-428232, T. urartu {10811}.

Gli-A2bs

Synonym	Gli-A[u] 2-XXI {10811}
dv	PI-428188, T. urartu {10811}.

Gli-A2bt

Synonym	Gli-A[u] 2-XXII {10811}
dv	PI-428244, T. urartu {10811}.

Gli-A2bu

Synonym	Gli-A[u] 2-XXIII {10811}
dv	PI-538733, T. urartu {10811}.

Gli-A2bv

Synonym	Gli-A[u] 2-XXIV {10811}
dv	PI-428212, T. urartu {10811}.

Gli-A2bw

Synonym	Gli-A[u] 2-XXV {10811}
dv	TRI-6734, T. urartu {10811}.

Gli-A2bx

Synonym	Gli-A[u] 2-XXVI {10811}
dv	PI-428254, T. urartu {10811}.

GLI-B2

Synonym	Gld 6B {1415}
Chromosome	6BS
Chromosome	6B
Varieties	CS.

Gli-B2a

Varieties

CS{988}.

Gli-B2b

Varieties

Bezostaya 1 {988}; Cobra {991}; Gladio {9986}; Sideral {991}.

Gli-B2c

Varieties

Courtot {991}; Escuala {9985}; Gabo {988}; Loreto {9986}; Manital {9986}; Prinqual {991}; Siete Cerros 66 {988}; Sinton {995};

Gli-B2d

Varieties

Akmolinka 1 {988}; Cesar {9981}; Friedland {991}; Tselinnaya 20 {988}.

Gli-B2e

Varieties

Arsenal {991}; Veronese {9986}; Zlatna Dolina {994}.

Gli-B2f

Varieties

Basalt {9981}; Maris Freeman {988}; Master {991}.

Gli-B2g

Varieties

Capitole {991}; Capelle-Desprez {991}; Galahad {988}; Forlani {9986}.

Gli-B2h

Varieties

Castan {991}; Mentana {9986}; Pane 247 {9985}; Partizanka {994}; Sadovo 1 {988}; Sistar{9986}.

Gli-B2i

Varieties

Insignia{988}; Robin{9981}.

Gli-B2j

Varieties

Farnese {9986}; Funo R250 {9986}; Novosadska Rana 1 {994}.

Gli-B2k

Varieties

Skala {988}.

Gli-B2l

Varieties

Clement {991}; Longbow {988}; Tracy {991}.

Gli-B2m

Varieties

Mironovskaya 808{988}; Open {991}; Renan{991}.

Gli-B2n

Varieties

Japhet {9981}; Rouge de Bordeau {9981}; Solo {988}.

Gli-B2o

Varieties

Hardi {9981}; Mara {9986}; Odesskaya 16 {988}; Pippo {9986}; Rivoli {991}; Slavjanka {9981}.

Gli-B2p

Varieties

Pliska {983}; Champtal {991}; Oderzo {9986}; Recital {991}; Gazul {9985}.

Gli-B2q

Varieties

Saratovskaya 39 {988}.

Gli-B2r

Varieties

Arminda {991}; Estica {991}; Genial {991}.

Gli-B2s

Varieties

Aquila {9981}; Saratovskaya 36 {988}.

Gli-B2t

Varieties

Tselinogradka {988}.

Gli-B2u

Varieties

Kirgizskaya Yubileinaya {988}.

Gli-B2v

Varieties

Declic {991}; Garant {991}; Libellula {9986}; Mahissa 1 {9985}; Poljarka {988}.

Gli-B2w

Varieties

Palata {9986}; Pembina {995}; Rieti DIV {9986}.

Gli-B2x

Varieties

Super Zlatna (biotype) {994}; Prostor {9981}; 251/83 {9981}.

Gli-B2y

Varieties

Centauro {9986}; E. Morandi {9986}.

Gli-B2z

Varieties

Maestro {9985}.

Gli-B2aa

Varieties

Salmone {9986}; E. Mottin {9981}.

Gli-B2ab

Varieties

Bordier {9981}; Orepi {991}.

Gli-B2ac

Varieties

Scipion {991}; Artaban {991}; Riol{991}; Lontra{9981}.

Gli-B2ad

Varieties

Champion {991}; Chopin {991}.

Gli-B2ae

Varieties

Priam {991}; Etoile d'Choisy {991}; Campeador {9985}; Krajinka (biotype) {994}.

Gli-B2af

Varieties

Montjuich {9985}; Mocho Sobarriba {9985}.

Gli-B2ag

Varieties

Jeja del Pais {9985}; Barbilla de Leon (MCB-1292) {9981}.

Gli-B2ah

Varieties

Rojo de Humanes (MCB-1262) {9981}; Grano de Miracolo {9981}.

Gli-B2ai

Varieties

Blanquillo (MCB-0908) {9981}.

Gli-B2aj

Varieties

Negrete de Malaga (MCB-1754) {9981}.

Gli-B2ak

Varieties

HY320 {9981}; Leader {9981}.

Gli-B2al

Varieties

Dankowska {991}.

Gli-B2am

Varieties

TM-275 {9981}; Uralochka {9981}.

Gli-B2an

Varieties

Eagle {9981}; Glenwari {9981}.

Gli-B2ao

Varieties

Olympic {9981}; Mokoan {9981}.

Gli-B2ap

Varieties

Veda {9981}; Magnif 27 {9981}.

Gli-B2aq

Varieties

Winglen {9981}; Isis {9981}.

Gli-B2ar

Varieties

Arbon {9981}; Roazon {9981}.

Gli-B2as

Varieties

Strela {9981}; Sredneuralskaya {9981}.

Gli-B2at

Varieties

Ranee {9981}; Javelin 48 {9981}.

Gli-B2au

Note	Null allele.
Varieties	Saratovskaya 29 {9987}.

GLI-D2

Synonym	Gld 6D {1415}
Chromosome	6DS
Chromosome	6D
Varieties	CS.

Gli-D2a

Varieties

CS {988}; Maris Freeman {988}; Sistar{9986}; Tracy {991}.

Gli-D2b

Varieties

Bezostaya 1 {988}; Cobra {991}; Farnese {9986}; Partizanka {994}.

Gli-D2c

Varieties

Escualo {9985}; Eridano {9986}; Rieti DIV {9986}; Siete Cerros 66 {988}.

Gli-D2d

Varieties

Dneprovskaya 521 {988}.

Gli-D2e

Varieties

Dollar {9985}; Lada {9981}; Mironovskaya 808 {988}; Open {991}.

Gli-D2f

Varieties

Creneau {991}; Kirgizskaya Yubileinaya {988}; Rempart{991}.

Gli-D2g

Varieties

Capelle-Desprez {991}; Futur {991}; Galahad {988}; Ghurka {988}; Mec {9986}.

Gli-D2h

Varieties

Capitole {991}; Chinook {995}; Eagle {00119}; Garant {991}; Sadovo 1 {988}; Thatcher {995}.

Gli-D2i

Varieties

Insignia 49 {00119}; Lario {9986}.

Gli-D2j

Varieties

Arcane {991}; Gallo {9986}; Gazul {9985}; Inia 66 {9985}; Mentana {9986}.

Gli-D2k

Varieties

Crvencapa {944}; Kzyl-Bas {988}; Skala {988}. Gli-D2l. Omitted. No reliable differences compared to existing alleles {9981}.

Gli-D2m

Varieties

Marquis {988}; Rex {991}; Rinconada {9985}; Suneca {119}; Veronese {9986}; Yecora 70 {9985}.

Gli-D2n

Varieties

Castan {991}; Champlein {991}; Mahissa 1 {9985}; Mercia {988}; Pippo {9986}.

Gli-D2o

Varieties

Omskaya 12 {988}. Cultivars Salmone and Resistente, which carry Gli-D2aa {9981}, were erroneously given as standards for allele Gli-D2o in {9986}.

Gli-D2p

Varieties

New Pusa {988}.

Gli-D2q

Varieties

Cook {9981}; E. Mottin {9981}; Fournil {991}; Volshebnitsa (biotype) {988}; Winglen {9981}; Soissons {991}.

Gli-D2r

Varieties

Kremena {988}; Mara {9986}; Montcada {9985}.

Gli-D2s

Varieties

Akmolinka 1 {988}; Bezenchukskaya 98 {988}; Selkirk (biotype) {995}.

Gli-D2t

Varieties

Golia {9986}; Gabo {9981}; Manital {9986}; Bokal {9981}.

Gli-D2u

Varieties

Loreto {9986}; Martial {991}; Cibalka {9981}.

Gli-D2v

Varieties

Epiroux {991}; Arbon {991}.

Gli-D2w

Varieties

Navarro 150 {9985}; Javelin {9981}; Hopps {9981}; Canaleja {9985}.

Gli-D2x

Varieties

Montjuich {9985}; Blanquillo{9985}.

Gli-D2y

Varieties

Candeal Alcala {9985}.

Gli-D2z

Varieties

Aragon 03 {9985}.

Gli-D2aa

Varieties

Salmone {9981}; Resistente {9981}.

Gli-D2ab

Varieties

Rojo de Boadilla de Campos (MCB-1031) {9981}.

Gli-D2ac

Varieties

Albatros {9981}.

Gli-D2ad

Varieties

Hembrilla Soria (MCB-1298) {9981}.

Gli-D2ae

Note	Null allele.
Varieties	Saratovskaya 29 (mutant) {9987

GLI-2 alleles were determined in 57 Yugoslav wheat varieties {994}.

GLI-Ag[i] 2

Note	6Ag[i ] {374}.
ad	Vilmorin 27/ Th. intermedium .

GLI-R

Gli-R2a

Note	d1 {03116}.
Varieties	Carnac hexaploid triticale {03116}.

Gli-R2b

Note	d2 {03116}.
Varieties	Mostral hexaploid triticale {03116}.

Gli-R2c

Note	t1 {03116}.
Varieties	Alamo hexaploid triticale {03116}.

Gli-R2d

Note	Null {03116}.
Varieties	Triticor hexaploid triticale {03116}.

Gli-R2e

Note	t2 {03115}.
Varieties	Tornado hexaploid triticale {03115}.

GLI-R[m] 2

Chromosome	6R
ad	CS/ S. montanum . The location of Gli-R2 in S. cereale is thought to have evolved from S. montanum {1339} via a translocation between 2R

GLI-S[l] 2

Chromosome

ad,su: CS/ Ae. longissima .

GLI-U2

Chromosome	6U
ad	CS/ Ae. umbellulata

GLI-V2

Chromosome	6VS
ad	Creso/ D. villosum .

GLI-3

GLI-A3

Synonym	Gld-2-1A {1416}
Chromosome	1AS
Varieties	Bezostaya 1. Each of the following GLI-A3 alleles, apart from Gli-A3d , which is a null, controls one minor omegagliadin with molecular mass about 41k

Gli-A3a

Varieties

CS, Prinqual, Courtot, Tselinogradka, Bezenchukskaya 98.

Gli-A3b

Varieties

Bezostaya 1.

Gli-A3c

Varieties

Anda.

Gli-A3d

Note	Null {9983}.
Varieties	Saratovskaya 210, Kharkovskaya 6, Richelle.

GLI-B3 {1119}, {422}. [ Glu-B2 {589}, Gld-B6 {422}]. 1BS {589}, {422}.

Gli-B3b

Varieties	Sicco. Gli-B3c {1119}, {422}.
Sources / synonyms	CS[*] /Thatcher1B.

GLI-R3

Chromosome	1RS
al	Four inbred lines (R2, J14, 8t, E2666).

GLI-S[l] 3

Note	1S[1] S {1228}. ad,su: CS/ Ae. longissima .
Marker associations	In Ae. longissima 2/Ae. longissima 10 , three gliadin loci, one glucose phosphate isomerase, and two glutenin loci were mapped relative to one another {1228}

Gli-V3

Chromosome	4VL
ad	Creso/ D. villosum .

GLI-5

GLI-A5

Chromosome	1AS
Varieties	Salmone.

Gli-A5a

Note	Null {9983}.
Varieties	CS.

Gli-A5b

Varieties

Marquis.

Allele Gli-A5b controls two slow-moving, easily-recognizable omega-gliadins. It is present in all common wheat cultivars having alleles Gli-A1m and Gli-A1r (and, probably, in those having Gli-A1e , Gli-A1l and Gli-A1q ), because earlier (for example, in {988}) two minor omega-gliadins encoded by Gli-A5b were considered controlled by these GLI-A1 alleles {9983}

GLI-B5

Chromosome	1BS
Varieties	Salmone.

Gli-B5a

Varieties

CS.

Gli-B5b

Varieties

Salmone.

In {988}, omega-gliadins controlled by GLI-B5 (allele Gli-B5b ) were attributed to alleles at the GLI-B1 locus (alleles Gli-B1c, i, k, m, n and o ).

GLI-6

Gli-A6a

Note	Null {9983}.
Varieties	CS; Bezostaya 1.

Gli-A6b

Varieties

Bezenchukskaya 98.

Gli-A6c

Varieties

Courtot, Anda, Mironovskaya 808.

GLI-7

GLI-A7

Chromosome	1DS
dv	AUS18913 {10547}.

The gamma-gliadin encoded by this locus co-segregated with the T1 omega-gliadin encoded by the GLID[t] T1 locus (currently included in the Catalogue as locus ( GLI-DT1 ). GLI-A7 was located 0.69 cM from GLI-D[t] 1 {10547}.

Inhibitors (dimeric) of heterologous alpha-amylase

IHA-B1.1

Chromosome	3BS
Varieties	CS {1260}.

IHA-B1.2 .

Iha-B1.2

Varieties

CS {0124}.

Iha-B1.2a

Chromosome	3BS
Varieties	CS {0124, 0125}.

Iha-B1.2b

Note	Null allele.
Varieties	Cadoux {0125}; Cranbrook {0125}; Tasman {0125}.

IHA-D1

Chromosome	3DS
Varieties	CS {1260}.

Subtilisin inhibition

SI-1 SI-R1

Chromosome	2RS
Chromosome	2R
ad	CS/Imperial, Holdfast/King II.

SI-H1

Synonym	Isa 1 {528}
Chromosome	2H
ad	CS/Betzes.

SI-2

SI-B2

Chromosome	1BS
su	Bersee (Koga II).

SI-D2

Chromosome	1DS
Varieties	Koga II.

SI-H2

Synonym	Ica 2 {528}, Ica 1 {528}
Chromosome	1H
ad	CS/Betzes.

SI-R2

Chromosome	1RS
Chromosome	1R
ad	CS/Imperial {529}.
tr	Gabo 1BL.1RS {701}.

SI-S[l] 2

Chromosome	1S
ad	CS/ Ae. longissima .

SI-U2

Chromosome	1U
ad	CS/ Ae. umbellulata . Considerable genetic variation for Si-2 was noted in {701}. A chromosome location for Si-H2 on 1HL was inferred in {528} but

Genes encoding proteins which inhibit the action of mammalian and insect, but not cereal, a-amylases, were located in chromosomes 3BS, 3DS and 6DS of Chinese Spring {1260}. Also, genes encoding inhibitors of insect a-amylases were in H. chilense chromosomes 4H[ch] and 7H[ch] {1262}.

Chromosome	1BL
Varieties	Recital{10908}.
Marker associations	Glu-B1 - 1.3 cM - Spa-B1 {10909}.

2. Proteins⌂ Home

2.1. Grain protein content⌂ Home

GPC-B1

2.2. Enzymes⌂ Home

ACPH-1

ACPH-2

AMP-2

AMP-3

a-AMY-1

a-AMY-A1

a-AMY-B1

a-AMY-2

b-AMY-1

EP-2

EST-1

EST-2

EST-3

EST-4

EST-7

LPX-1

LPX-2

AADH-1

NDH-2

AHASL 1

PSY-1

PPO-2

2.3. Endosperm storage proteins⌂ Home

GLU-2

GLU-3

GLU-4

GLU-5

GLI-1

GLI-2

GLI-R

GLI-3

GLI-5

GLI-6

GLI-7

Inhibitors (dimeric) of heterologous alpha-amylase

Subtilisin inhibition

TI-1

TI-2

SPA-1